principles of systematics and nomenclature general system and phylogeny of insects systematics of Ephemeroptera

last update 1.IX.2020


Type specimens of mayflies (Ephemeroptera) in the
Zoological Institute
of Russian Academy of Sciences
(in Saint-Petersburg, Russia)

By N.J. Kluge

For details about old type-specimens (original labels, some morphological characters, comments on lectotype designation and synonymy) see:

.. 1995. . (Ephemeroptera). . . . , .: 52 . PDF

Kluge N.J. 1995. A catalogue of the type-specimens in the collection of the Zoological Institute, Russian Academy of Sciences. Insecta, Ephemeroptera. Zool. Inst. Rus. Acad. Sci., St.Petersburg: 52 pp. (in Russian).

In 19931994 all type specimens of extant mayflies formerly deposited in Moscow State University and Paleontological Institute (Moscow) were transferred to Zoological Institute (S.-Petersburg). Now type collection of the Zoological Institute includes deposited type specimens of extant Ephemeroptera species described by A.D. Hummel (1825), L. Navas (1912), P. Esben-Petersen (1916), O.A. Tshernova and her co-authors (19281985), K.A. Brodsky (1930), A.T. Buldovsky (1935), R.S. Kazlauskas (19591964), O.Ya. Bajkova (19621980), N.D. Sinitshenkova and her co-authors (since 1973 and later), L.A. Kustareva (19761984), V.V. Belov (1979, 1981), N.J. Kluge and his co-authors (since 1979 and later), E.A. Novikova and her co-authors (since 1987 and later) and some others. Some type specimens are being temporally held on the premises of the Department of Entomology of St. Petersburg State University.

Type-specimens of mayflies described by T.M. Tiunova and I.M. Levanidova (since 1989 and later) are deposited in the Institute of Biology and Soil Science (Vladivostok). 

All type-specimens of fossil Ephemeroptera described in the USSR and Russia are deposited in the Paleontological Institute (Moscow).

       

Abbreviations: 
ZIN Zoological Institute (S-Petersburg) 
L-S-I imago reared from larva, with larval and subimaginal exuvia; 
L-S subimago reared from larva, with larval exuvia;
L/S mature larva with extracted parts of subimago;
link to the catalogue Ephemeroptera of the World.

    

Arbitrary signs

   

Numbers of specimens collected by N.J. Kluge

* * *

abnormis Tshernova 1949: 139 [Heptagenia
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

abracadabrus Kluge 1983: 31 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

acinaciger Kluge 1983: 70 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

adjaricus Kluge & Godunko & Apanaskevich 2013 [Ecdyonurus (Helvetoraerticus)]
Type deposition. ZIN: holotypus (imago ) & paratypi

albicauda Kluge 1987: 318 [Heptagenia (Heptagenia) sulphurea, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi 

albisternum Novikova (in Tshernova et al.) 1986: 131 [Cloeoptilum
Type deposition.
ZIN: holotypus (L-S-I) & paratypi (see Novikova 1987)

alboniger Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

album Kluge 2016 [Procloeon (Oculogaster)]
Type deposition. ZIN: holotypus (L/S) & paratypi

alcarrazae Kluge 1992: 18 [Baetis (Caribaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

alexandrae Brodsky 1930: 697 [Ameletus
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

alexandri Kluge & Tiunova 1989: 12 [Epeorus (Iron)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi 

altaica Tshernova 1949: 142 [Cinygmula
Type deposition.
ZIN: syntypi (larvae)

altaicus Kluge 2007: 251 [Ameletus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

altioculus Kluge 1984: 722 [Choroterpes (Euthraulus)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

amazonicus Kluge 2009 [Terpides (Fittkaulus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ameliae Kluge & Gattolliat & Salles 2017 [Crassabwa]
Type deposition. ZIN: holotypus ( larva and extracted subimago) & paratypi (part)

amurensis Kluge (in Tshernova et al.) 1986: 141 [Caenis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi (see Kluge 1987)

amurensis Navas 1912: 414 [Ephemera
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypus

anatolii Sinitshenkova 1981: 814 [Epeorus (s.str.)] 
Type deposition.
ZIN: holotypus (larva) & paratypus

andrianovae Tshernova & Belov 1982: 283 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

arlecchino Kluge 2007: 254 [Ameletus montanus, subsp.]
!!  Type deposition. ZIN: holotypus (L-S-I) & paratypi

arsenjevi Tshernova 1952: 255 [Heptagenia
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

asiatica Tshernova & Bajkova 1960: 413 [Protobehningia
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

asiatica Sinitshenkova 1973: 13 [Rhithrogena
!! Type deposition. ZIN: holotypus (larva) & paratypi

aspersus Kluge 1980: 569 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

aurarius Kluge 1983: 29 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

bacillus Kluge 1983: 72 [Baetis gracilis, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

baconaoi Kluge 1994: 276 [Hagenulus (Careospina)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

bajkovae Kluge (in Tshernova et al.) 1986: 117 [Ecdyonurus
Objective synonym
: zhiltzovae Bajkova 1975 [Paracinygmula]

balticus Kazlauskas & Sanvaitite 1962: 41 [Baetopus
TYPES LOST

barnardi Kluge 2020 [Procloeon (Oculogaster)]
Holotype deposition. Albany Museum in South Africa (L-S-I).
Paratypes deposition. ZIN.

binerve Kluge 1987: 775 [Rhithrogena
!! Type deposition. ZIN: holotypus (L-S-I) & paratypi

binotata Sinitshenkova 1982: 65 [Rhithrogena
Type deposition. ZIN: holotypus ( imago) & paratypi

bispinosa Kluge & Novikova 2011 [Acentrella (Leibebiella)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

bipunctata Esben-Petersen 1916: 7 [Heptagenia
TYPES LOST

bogori Kluge 2014 [Choroterpes (Dilatognathus)]
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

borysthenica Tshernova 1937: 5 [Oligoneuriella
Type deposition.
ZIN: syntypi (larvae)

brodskyi Kustareva 1976: 66 [Rhithrogena
Type deposition.
ZIN: holotypus (parts of imago) & paratypus

brodskyi Bajkova 1979: 308 [Siphlonurus
Type deposition.
ZIN: holotypus (L-S)

carnivora Kluge 2015 [Rhithrogena (Himalogena)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

cataractae Kluge 2012 [Choroterpes (Dilatognathus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

caucasica Tshernova 1938: 58 [Cinygma
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

caucasica Tshernova 1931: 216 [Habroleptoides
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

caucasicus Kluge 2012 [Choroterpes (Euthraulus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

cedrensis Sinitshenkova 1977: 44 [Ameletus
Type deposition.
ZIN: holotypus ( imago) & paratypi

celebensis Kluge 2010 [Tricorythus (Sparsorythus)]
Type deposition. ZIN: holotypus (male L-S-I) & paratypi

chankae Tshernova 1952: 263 [Siphlonurus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1985) & paralectotypus

chantauensis Kluge 1981: 1418 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

chimaera Kluge 2008: 398 [Hermanella (Hylister)]
Type deposition. ZIN: holotypus (male L-S-I) & paratypi

citrina Hummel 1825: 71 [Ephemera] (nomen oblitum) 
Type deposition.
ZIN: lectotypus ( subimago, designated by Kluge 1995)

clavifolium Kluge 2016 [Cheleocloeon]  
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

confinis Tshernova 1928: 322 [Ecdyonurus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

contamanensis Kluge 2015 [Terpides]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

continentale Kluge & Suttinun 2020 [Indocloeon (Indocloeon)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

corniger Kluge 1991: 16 [Brachycercus
Type deposition.
ZIN: holotypus (L-S)

cornuta Tshernova 1952: 286 [Ordella
Type deposition.
ZIN: lectotypus ( larva, designated by Kluge 1995) & paralectotypi

cothurnata Tshernova 1952: 267 [Paraleptophlebia
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

cristatus Bajkova 1976: 586 [Ameletus
TYPES LOST

cubensis Malzacher & Naranjo & Gonzalez-Lazo & Kluge 2007 [Caenis]
!! Type deposition. ZIN: holotypus (L-S-I) & paratypi

cubensis Kluge & Naranjo 1990: 576 [Tricorythodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

cylindroculata Kluge & Novikova 2011 [Acentrella (Leibebiella)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

daedaleus Sinitshenkova 1981: 816 [Epeorus (Belovius)] 
Type deposition.
ZIN: holotypus (larva) & paratypi

decolorata Sinitshenkova 1973: 16 [Rhithrogena
Type deposition.
ZIN: holotypus ( imago) & paratypi

dentata Bajkova 1967: 327 [Ephemerella
Type deposition.
ZIN: holotypus (larva)

depapricus Kluge 2013 [Nonnullidens]
Type deposition. ZIN: holotypus (L

desertus Novikova & Kluge 1987: 14 [Baetis (Labiobaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

diptera Kluge & Novikova 2011 [Acentrella]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

diversicolor Tshernova 1952: 266 [Baetis
Type deposition. ZIN: holotypus (larva) (Kluge 2012)

dracon Kluge 1983: 27 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

eatoni Brodsky 1930: 33 [Ametropus]
Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypus

echinovaris Kluge 2015 [Terpides]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

edentatum Kluge 2019 [Potamocloeon]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

elgonensis Kluge 2012 [Rhithrocloeon (Kivuiops)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ermolenkoi Tshernova 1981: 327 [Epeorus (Belovius)] 
Type deposition.
ZIN: holotypus ( imago) & paratypi

eugeniae Kluge 1983: 1116 [Rhithrogena
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

eugenii Sinitshenkova & Varychanova 1989: 576 [Ameletus
Type deposition.
ZIN: holotypus ( imago) & paratypi

europaeus Kluge 1991: 19 [Brachycercus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

evanescens Kluge 1994: 276 [Hagenulus (Careospina)] 
Type deposition.
ZIN: holotypus ( imago) & paratypi

excisa Sinitshenkova 1979: 7 [Rhithrogena
Type deposition.
ZIN: holotypus ( imago) & paratypi

exophthalmus Kluge 2010 [Tricorythus]
Type deposition. ZIN: holotypus (L-S) & paratypi

fascipennis Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

feles Kluge 1980: 562 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

femorata Tshernova 1972: 611 [Asiatella
Type deposition.
ZIN: holotypus (larva)

fenestratum Kazlauskas 1963: 588 [Pseudocloeon
Type deposition.
ZIN: holotypus (parts of larva)

flavus Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

frater Tshernova 1938: 58 [Ecdyonurus
Type deposition.
ZIN: holotypus ( imago)

frolenkoi Sinitshenkova 1981: 815 [Epeorus (Belovius)] 
Type deposition.
ZIN: holotypus (larva) & paratypi

furcifer Kluge 2016 [Tricorythus]
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

fusca Kluge & Novikova 2016: 28 [Pseudopannota
Type deposition.
ZIN: holotypus (L/S) & paratypi

fusconotum Kluge 2009 [Terpides (Tikuna)]  
Type deposition.
ZIN: holotypus (S-I) & paratypi

fuscus Sinitshenkova 1976: 859 [Iron
Type deposition.
ZIN: holotypus (larva) & paratypi

ganeshi Kluge 2014: 525 [Notophlebia]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

gilliesi Kluge & Novikova 2017: 52 [Ephemerythus]
Type deposition. ZIN: holotypus ( larva and extracted subimago with eggs)

gnom Kluge 1983: 74 [Pseudocloeon
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

gornostajevi Tshernova 1981: 327 [Epeorus (Belovius)] 
?? Type deposition. ZIN: holotypus ( imago) & paratypi

gracilis Tshernova 1952: 257 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

grallator Kluge & Naranjo 1990: 575 [Tricorythodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

grandifolia Tshernova 1952: 256 [Cinygmula
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

grandis Kluge 1994: 265 [Hagenulus (Turquinophlebia)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

grisea Navas 1912: 416 [Andromina
Type deposition. ZIN: syntypi ( subimagines) 

grunini Sinitshenkova 1982: 59 [Iron
Type deposition.
ZIN: holotypus ( imago) & paratypi

gultsha Kustareva 1984: 16 [Iron
TYPE LOST

gunti Kluge 2015 [Rhithrogena (Himalogena)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

guranica Belov 1981: 601 [Heptagenia
Type deposition.
ZIN: holotypus ( imago) & paratypi

gurvitshi Kustareva 1984: 20 [Iron
TYPES LOST

heptapotamicus Brodsky 1930: 693 [Baetis
Type deposition.
ZIN: holotypus ( imago)

heterophillum Kluge & Novikova 1992: 74 [Cloeon (Procloeon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

hissari Kluge 1985: 1586 [Caenis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

hissaricus Novikova 1991: 793 [Baetis ursinus, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

hyalinus Esben-Petersen 1916: 6 [Ecdyonurus] nom. praeocc. 
Type deposition.
ZIN: holotypus ( imago) 
Objective synonym
: peterseni Lestage 1930 [Ecdyonurus]

imanica Bajkova 1972: 207 [Rhithrogena
Type deposition.
ZIN: holotypus ( imago)

immanis Kluge 1985: 18 [Siphlonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

inaccessibile Kluge & Naranjo 1994: 778 [Euthyplocia
Type deposition.
ZIN: holotypus ( imago) & paratypi

indonesiae Kluge 2011 [Indocloeon]
Type deposition. ZIN: holotypus (L-S-I) & paratypi  

inexpectatum Tshernova 1928: 320 [Pseudocloeon
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

inferior Kluge 1991: 133 [Cloeodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

ingridae Kluge & Godunko & Svitok 2020 [Centroptella]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

intermedia Brodsky 1930: 708 [Iron montanus, var.] 
Type deposition.
ZIN: syntypi (imagines)

inversus Kluge 1980: 569 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

irina Tshernova & Belov 1982: 286 [Cinygmula
Type deposition.
ZIN: paratypus ( imago); holotypus lost

issyksuvensis Brodsky 1930: 688 [Baetis
Type deposition.
ZIN: syntypi

itayana Kluge 2007: 130 [Oligineuria]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

karasuensis Kustareva 1976: 58 [Drunella
Type deposition.
ZIN: holotypus (larva)

kazlauskasi Kluge 1983: 78 [Centroptilum
  Type deposition.
ZIN: holotypus (L-S-I) & paratypi

kirgisicus Kustareva 1984: 14 [Iron
TYPES LOST

kogistani Novikova & Kluge 1994: 637 [Baetis (Nigrobaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

kopetdagi Kluge 1985: 1586 [Caenis
  Type deposition.
ZIN: holotypus ( L-S-I) & paratypi

kozhovi Bajkova 1967: 327 [Ephemerella
Type deposition.
ZIN: holotypus ( imago) & paratypi

kurenzovi Bajkova 1965: 301 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

labiatus Sinitshenkova 1981: 75 [Ameletus
Type deposition.
ZIN: holotypus (larva) & paratypi 

laciniosa Sinitshenkova 1979: 816 [Rhithrogena
Type deposition.
ZIN: holotypus (larva) & paratypi

lancetofolium Kluge 2016 [Cheleocloeon]
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

latericius Sinitshenkova 1982: 53 [Iron
Type deposition.
ZIN: holotypus ( imago) & paratypi

latifolia Kluge 2014 [Nousia (Araucophlebia)] 
Type deposition. ZIN: holotypus ( L-S-I) & paratypi

latifrons Tshernova & Belov 1982: 287 [Cinygmula
?? Type deposition. ZIN: paratypi; holotypus lost

latipes Tshernova 1952: 273 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

latipennis Kluge & Bernal 2018 [Moribaetis]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

lenoki Tshernova 1952: 275 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

lepnevae Tshernova 1949: 152 [Ephemerella 
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

lepnevae Brodsky 1930: 39 [Rhithrogena
Type deposition.
ZIN: syntypi ( imagines)

levanidovae Tshernova 1952: 274 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

levanidovae Sinitshenkova 1982: 60 [Iron
Type deposition.
ZIN: holotypus ( imago) & paratypi

levanidovi Tshernova & Belov 1982: 289 [Cinygmula
Type deposition.
ZIN: holotypus ( imago)

levis Navas 1912: 415 [Epeorus
Type deposition. ZIN: holotypus ( imago) 

lilliputian Kluge 1991: 134 [Paracloeodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypus 

lithuanica Kazlauskas 1959: 168 [Ephemerella
TYPES LOST

longifolius Kluge 1992: 44 [Baetis (Fallceon)] 
Type deposition.
ZIN: holotypus (male L-S-I) & paratypi

longilobata Tshernova 1928: 113 [Leptophlebia (Paraleptophlebia)] 
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

longipes Tshernova 1952: 274 [Ephemerella
Type deposition.
ZIN: holotypus (larva)

longulus Sinitshenkova 1981: 77 [Ameletus
Type deposition.
ZIN: holotypus (larva) & paratypi

ludmilae Kluge 2012 [Choroterpes (Choroterpes)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ludmilae Kluge & Gattolliat & Salles 2017 [Crassabwa]
Type deposition. ZIN: holotypus (L-S-I) & paratypi (part)

ludmilae Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

lunata Tshernova 1952: 269 [Paraleptophlebia
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

lunamontana Kluge & Novikova 2011 [Acentrella (Tanzaniops)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

lutea Kluge 1987: 311 [Heptagenia (Heptagenia)] nom. praeocc. 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

macronyx Kluge in Tshernova et al. 1986: 141 [Caenis
Type deposition.
ZIN: holotypus (mature larva) & paratypi

macronyx Kluge & Novikova 1992: 76 [Cloeon (Procloeon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

macronyx Kluge 1996 [Metreplecton]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

maculata Tshernova1952: 287 [Ordella
Type deposition.
ZIN: syntypi (larvae)

maculatus Tshernova 1949: 148 [Iron
Type deposition.
ZIN: lectotypus (larva, designated by Kluge & Tiunova 1989) & paralectotypi

magnus Tshernova 1952: 284 [Brachycercus
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1991) & paralectotypi

malabarensis Kluge 2020 [Procloeon (Oculogaster)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

mamasae Kluge & Novikova 2011 [Platybaetis]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

maritimus Kluge 1983: 76 [Centroptilum
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

markevitshi Belov 1979: 577 [Uracanthella
Type deposition.
ZIN: holotypus (larva) & paratypi

mercatorius Kluge 2012 [Choroterpes (Choroterpes)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

merga Peters & Gillies 1991 [Protobehningia]
Type deposition.
ZIN: paratypi

micracantha Kluge 1989: 4 [Heptagenia (Dacnogenia) coerulans, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

microfolius Kluge & Novikova 2014 [Indobaetis]
Type deposition. ZIN: holotypus (L-S) & paratypi

micus Bajkova 1976: 583 [Ameletus
Type deposition.
ZIN: holotypus (larva)

miliaria Tshernova 1952: 286 [Ordella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

minima Sinitshenkova 1973: 13 [Rhithrogena
Type deposition. ZIN: holotypus (larva) & paratypi

minuta Brodsky 1930: 709 [Iron montanus, var.] 
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

minutus Tshernova 1952: 285 [Brachycercus
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1991) & paralectotypi

modesta Brodsky 1930: 31 [Ephemera
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

mongolica Bajkova & Varychanova 1978: 14 [Heptagenia
TYPES LOST

mongolicus Tshernova 1952: 266 [Baetis
Type deposition. ZIN: lectotypus and paralectotypi (larvae) (Kluge 2012)

monilistylus Kluge & Tiunova & Novikova 2014 [Procloeon]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

monophyllus Kluge 2012 [Choroterpes (Monophyllus)]  
Type deposition. ZIN: holotypus (L) & paratypi

montana Brodsky 1930: 704 [Iron montanus, var.] 
Type deposition.
ZIN: syntypi

montanus Kluge & Naranjo 1990: 574 [Tricorythodes
 Type deposition.
ZIN: holotypus (L-S-I) & paratypi

munyagae Kluge 2012 [Rhithrocloeon (Kivuiops)]
Type deposition. ZIN: holotypus (L-S/I) & paratypi

mycetopis Brodsky 1930: 691 [Baetis
Type deposition.
ZIN: holotypus ( imago)

naranjoi Kluge 1992: 19 [Baetis (Americabaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

natans Buldovsky 1935: 153 [Chankagenesia
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

niger Kluge 2013 [Nonnullidens]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

niger Kluge 2020 [Procloeon (Oculogaster)]
Holotype deposition. Albany Museum in South Africa (L-S-I).
Paratypes deposition. ZIN.

niger Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

nigrabdominalis Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (S-I) & paratypi

nigricolor Kluge & Novikova 2017: 41 [Ephemerythus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

nigridorsum Tshernova 1934: 240 [Eopolymitarcys
Type deposition.
ZIN: syntypi (imagines)

nigripes Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L/S) & paratypi

nigripilosus Sinitshenkova 1976: 860 [Iron
Type deposition.
ZIN: holotypus (larva) & paratypi

nigrobulla Kluge 2009 [Terpides (Tikuna)]
Type deposition. ZIN: holotypus (L-S) & paratypi

nigrohumeris Kluge 2017: 114 [Cloeodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

nigromaculata Brodsky 1930: 708 [Iron montanus, var.] 
Type deposition.
ZIN: syntypi

nigrotibialis Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

nikolskyi Tshernova 1948: 1453 [Acanthametropus
Type deposition.
ZIN: holotypus (larva)

ninae Kluge 1995 [Epeorus (Belovius)] 
Type deposition.
ZIN: holotypus ( imago) & paratypi 
Objective synonym: sinitshenkovae Tshernova 1981 [Epeorus (Belovius)]

nuda Tshernova 1949: 53 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

obscuriventris Tshernova 1952: 266 [Baetis
Type deposition. ZIN: lectotypus and paralectotypi (larvae) (Kluge 2012)

obtusiceps Tshernova 1952: 265 [Baetis
Type deposition. ZIN: lectotypus and paralectotypi (larvae) (Kluge 2012)

odontostylus Kluge & Novikova 2017: 463 [Anafroptilum
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

orbiticola Kluge 1987: 305 [Heptagenia (Kageronia)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

oreophila Kustareva 1978: 92 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

oreophilus Kluge 1982: 15 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

oriens Bae & McCafferty 1991: 54 [Potamanthus luteus, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I)

orientalis Kluge 1983: 74 [Baetis atrebatinus, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

orientalis Tshernova 1952: 279 [Ephemerella
!! Type deposition. ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

oriente Kluge 1994: 282 [Hagenulus (Traverina)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

ornata Tshernova 1972: 613 [Ephemerellina
Type deposition.
ZIN: holotypus ( subimago) & paratypus

ornatipennis Tshernova 1938: 55 [Ecdyonurus
Type deposition.
ZIN: holotypus ( imago)

ornatipennis Kluge 2020 [Procloeon (Monilistylus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ornatodermis Kluge 2015 [Terpides]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ornatum Tshernova 1928: 319 [Procloeon
Type depositions.
ZIN: paralectotypi; 
Zool. Mus. Hamburg Univ.: lectotypus ( imago), designated by Sowa 1975: 108

orthostylus Kluge & Novikova 2017: 461 [Anafroptilum
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

pacoi Kluge 1994: 269 [Hagenulus (Poecilophlebia)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

padunica Kazlauskas 1963: 584 [Ephemerella (Tortleya)] 
TYPES LOST

palaearcticus Tshernova 1931: 217 [Oniscigaster
Type deposition.
ZIN: holotypus ( imago)

pallidus Tshernova 1928: 114 [Brachycercus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1991) & paralectotypi

panamensis Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

pamirensis Kustareva 1984: 20 [Iron
TYPE LOST

pamirica Kluge 2015 [Rhithrogena (Himalogena)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

pannota Kluge & Novikova 2016: 11 [Pseudopannota
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

paradoxa Buldovsky 1935: 157 [Anagenesia
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paratypi

parvus Kluge 1979: 807 [Ameletus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

pellucida Brodsky 1930: 35 [Cinygma
Type deposition.
ZIN: holotypus ( imago)

perflava Brodsky 1930: 701 [Heptagenia
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1987) & paralectotypus

peterseni Lestage 1930: 199 [Ecdyonurus
Type deposition.
ZIN: holotypus ( imago)
Invalid objective synonym
: hyalinus Esben-Petersen 1916 [Ecdyonurus] nom. praeocc.

petropolitanum Kluge & Novikova 1992: 68 [Cloeon (Intercloeon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

petrovi Tshernova 1938: 55 [Baetis
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypus

pilatus Sinitshenkova 1981: 73 [Ameletus
Type deposition.
ZIN: holotypus (larva) & paratypi

planifrons Kluge 1992: 15 [Baetis (Caribaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

platysma Tshernova 1941: 239 [Heptagenia
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

plesius Kluge 2008: 394 [Simothraulopsis (Maculognathus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

polita Bajkova 1970: 146 [Isonychia
Type deposition.
ZIN: holotypus ( imago)

pontica Kluge 1984: 40 [Habroleptoides
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

procerus Bajkova 1976: 584 [Ameletus
Type deposition.
ZIN: holotypus (larval exuvia) & paratypi (see Kluge 1995)

pseudothermicus Kluge 1983: 65 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

putoranica Kluge 1980: 576 [Cinygmula
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

quadrata Kluge & Zhou (in Kluge & Zhou & Jacobus & McCafferty) 2004 [Ephemerella]
Type deposition. ZIN: holotypus (larva) & paratypi

quadrinotata Sinitshenkova 1982: 63 [Rhithrogena
Type deposition. ZIN: holotypus ( imago) & paratypi

quadripunctata Kluge 1989: 4 [Heptagenia (Heptagenia)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

rautiani Sinitshenkova 1982: 53 [Epeorus (Epeorus)] 
Type deposition.
ZIN: paratypi (larvae, subimageos, imageos); holotypus lost 

reductus Kluge 2013 [Nonnullidens]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

rheophilus Brodsky 1930: 712 [Iron
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

rohdendorfi Tshernova 1985: 15 [Rhoenanthus
Type deposition.
ZIN: holotypus ( imago)

romantzovi Kluge 1988: 76 [Ephemera
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

rossicus Kluge 2007: 253 [Ameletus montanus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

rossicus Tshernova 1928: 321 [Ecdyonurus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

rubrofasciatus Brodsky 1930: 716 [Ecdyonurus
Type deposition.
ZIN: holotypus ( imago)

sabalo Kluge 2008: 389 [Simothraulopsis (Maculognathus)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sacculobranchis Kluge & Naranjo 1990: 570 [Tricorythodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

scabriventris Kluge & Novikova 2011 [Acentrella]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

scalaris Kluge 1983: 34 [Ecdyonurus
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

secundum Kluge & Suttinun 2020 [Indocloeon (Indocloeon)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

semicarnivora Kluge 2015 [Rhithrogena (Himalogena)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

setigera Bajkova 1965: 327 [Ephemerella
Type deposition.
ZIN: holotypus (larva)

sextus Kluge 1992: 45 [Baetis (Fallceon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

sextus Kluge 1994: 261 [Hagenulus (Borinquena)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

shadini Kazlauskas 1964: 170 [Pseudocentroptilum
Type deposition.
ZIN: lectotypus (parts of larva, designated by Kluge 1995) & paralectotypus

sibirica Tshernova 1952: 278 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

sibirica Tiunova & Kluge & Ishiwata 2004 [Isonychia ussurica
Type deposition. ZIN: holotypus (L-S-I) and paratypi

sibirica Brodsky 1930: 37 [Rhithrogena
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

sibiricum Kazlauskas 1963: 589 [Pseudocloeon
Type deposition.
ZIN: holotypus (parts of larva) & paratypi

sierramaestrae Kluge 1994: 274 [Hagenulus (Careospina) hespera, subsp.] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

sierramaestrae Kluge & Naranjo 1990: 570 [Tricorythodes
 Type deposition.
ZIN: holotypus (L-S-I) & paratypi

silvaticus Kluge 1983: 67 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

sinitshenkovae Tshernova 1981: 332 [Epeorus (Belovius)] 
Type deposition.
ZIN: holotypus ( imago) & paratypi
Objective synonym
: ninae Kluge 1995 [Epeorus (Belovius)]

sinitshenkovae Tiunova & Kluge & Ishiwata 2004: 28 [Isonychia vshivkovae
Type deposition. ZIN: holotypus (L-S-I) and paratypi

smirnovi Tshernova 1978: 74 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

soldatovi Tshernova 1952: 253 [Heptagenia
Type deposition. ZIN: lectotypus (larva, designated by Kluge 1987) & paralectotypi

solida Bajkova 1980: 796 [Ephemerella
TYPES LOST

spiniventre Kluge & Novikova 1992: 65 [Cloeon (Intercloeon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

spinosa Tshernova & Belov 1982: 291 [Cinygmula
Type deposition.
ZIN: holotypus ( imago)

stackelbergi Sinitshenkova 1973: 18 [Rhithrogena
Type deposition.
ZIN: holotypus ( imago) & paratypus

stipposus Kluge 1982: 18 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

subcostalis Kluge 2012 [Choroterpes (Choroterpes) picteti]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

sublongicauda Tshernova 1949: 306 [Palingenia
Type deposition.
ZIN: syntypi

submontana Brodsky 1930: 683 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

sumbarensis Kluge 1984: 722 [Choroterpes (Euthraulus)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

superior Kluge 1991: 130 [Cloeodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

tadjikistanicus Brodsky 1930: 696 [Cloeon
Type deposition.
ZIN: syntypi ( imagines)

tadzhikorum Tshernova 1931: 218 [Heptagenia
Type deposition.
ZIN: holotypus ( imago)

taeniata Tshernova 1952: 279 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

talasi Novikova & Kluge 1994: 635 [Baetis (Nigrobaetis)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

taldybulaki Sroka & Godunko & Novikova & Kluge 2012 [Baetis (Rhodobaetis)]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

tenax Tshernova 1952: 273 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

tener Kluge 2016 [Tricorythus]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

testudineus Kluge 1992: 45 [Baetis (Fallceon)] 
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

thani Tshernova 1972: 609 [Vietnamella
Type deposition.
ZIN: holotypus (larva) & paratypus

thymalli Tshernova 1952: 277 [Ephemerella
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

tianshanica Kustareva 1984: 18 [Iron
TYPES LOST

tianshanica Brodsky 1930: 37 [Rhithrogena
Type deposition.
ZIN: lectotypus (L/S, designated by Kluge 1995) & paralectotypi

tianshanica Kustareva 1984: 26 [Stenonema
Type deposition.
ZIN: holotypus ( imago)

transbajkalica Tshernova 1973: 333 [Ephemera
Type deposition.
ZIN: holotypus ( imago) & paratypi

transiliensis Brodsky 1930: 689 [Baetis
Type deposition.
ZIN: holotypus ( imago)

triacantha Tshernova 1949: 41 [Ephemerella
TYPES LOST

tricolor Tshernova 1928: 320 [Baetis
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

truncifolium Kluge 2016 [Cheleocloeon]  
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

tshernovae Edmunds & Traver 1959 [Behningia
Type deposition.
ZIN: holotypus ( imago) & paratypi

tshernovae Bajkova 1962: 202 [Ephemerella
Type deposition.
ZIN: holotypus (larva)

tshernovae Sinitshenkova 1982: 56 [Iron
Type deposition. ZIN: holotypus ( imago) & paratypi

tshernovae Kazlauskas 1963: 582 [Neoephemera
Type deposition.
ZIN: holotypus (larva) & paratypi

tuberculatum Kazlauskas 1963: 586 [Pseudocloeon (Baetiella)] 
Type deposition.
ZIN: holotypus (parts of larva)

tubulatus Tshernova 1952: 285 [Brachycercus
Type deposition.
ZIN: holotypus (larva)

ugandanus Kluge 2010 [Ephemerythus niger]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

ulmeri Brodsky 1930: 686 [Caenis
Type deposition.
ZIN: syntypi ( imagoes)

unguiculatum Tshernova 1941: 242 [Cloeon
Type deposition.
ZIN: lectotypus (parts of larva, designated by Kluge 1996)

unicolor Tshernova 1952: 250 [Rhithrogena
Type deposition.
ZIN: lectotypus (L/S, designated by Kluge 1995) & paralectotypi

unicolorata Tshernova 1979: 9 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

ursinus Kazlauskas 1963: 585 [Baetis
Type deposition.
ZIN: holotypus (parts of larva)

ussurica Bajkova 1970: 146 [Isonychia
Type deposition.
ZIN: holotypus ( imago) & paratypus

ussuricus Kluge 1983a: 68 [Baetis
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

variegatus Kluge 2013 [Nonnullidens]
Type deposition. ZIN: holotypus (L-S-I) & paratypi

vernalis Tshernova 1928: 115 [Siphlonurus
Type deposition.
ZIN: holotypus (genitals of imago)

verrucosa Kluge 1980: 566 [Ephemerella
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

vibratorius Kluge 2017: 96 [Cloeodes
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

viviparus Kluge 2020 [Thraulodes]
Type deposition. ZIN: holotypus (S-I) & paratypi

vladivostokica Kluge 1982: 112 [Paraleptophlebia
Type deposition.
ZIN: holotypus (L-S-I) & paratypi

vshivkovae Tiunova & Kluge & Ishiwata 2004: 24 [Isonychia
Type deposition. ZIN: holotypus (L-S-I) and paratypi

werestschagini Tshernova 1952: 253 [Heptagenia
Type deposition.
ZIN: lectotypus (larva, designated by Kluge 1995) & paralectotypi

zachvatkini Tshernova 1952: 257 [Cinygmula
Type deposition.
ZIN: holotypus ( imago) & paratypi

zaitsevi Tshernova 1981: 326 [Epeorus (Epeorus)] 
Type deposition.
ZIN: holotypus ( imago)

zapekinae Bajkova 1967: 327 [Ephemerella
Type deposition.
ZIN: holotypus ( imago) & paratypi

zhelochovtsevi Tshernova 1952: 263 [Siphlonurus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1985a) & paralectotypus

zhiltzovae Tshernova 1972: 605 [Ecdyonurus
Type deposition.
ZIN: holotypus ( imago) & paratypi

zhiltzovae Bajkova 1975: 54 [Paracinygmula
Type deposition.
ZIN: holotypus (larva) & paratypi
Objective synonym
: bajkovae Kluge 1986 [Ecdyonurus]

zhiltzovae Sinitshenkova 1979: 811 [Rhithrogena
Type deposition.
ZIN: holotypus ( imago) & paratypi

zimini Tshernova 1931: 214 [Cloeon
Type deposition.
ZIN: holotypus ( imago) & paratypi

znojkoi Tshernova 1938: 56 [Ecdyonurus
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

znojkoi Tshernova 1938: 59 [Iron
Type deposition.
ZIN: lectotypus ( imago, designated by Kluge 1995) & paralectotypi

 to general index