CLADOENDESIS OF EPHEMEROPTERA

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Plesiomorphon Ephemerella/fg5

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Ephemerella/fg1
Tricorythus/fg1Ephemerella/fg2Timpanoga/fg1Ephemerella/fg3Torleya/g1Ephemerella/fg4 - pm.Ephemerella/fg5)

Nomen hierarchicum: Ephemerella/fg5 [f:1909; g:1862] (sine Drunella, Caudatella, Cincticostella)

In circumscription fits:

subgen. Ephemerella: Edmunds 1959: 544

gen. Ephemerella: Allen 1980: 74

plesiomorphon Ephemerella/fg5: Kluge 2004: 104

Nominal taxa included: 

Chitonophora/g [g:1908]

Caurinella/g [g:1984]

Zonadia/g [g:2008]

Hosoba/g [g:2008]

Draeconia/g [g:2008]

Scholitza/g [g:2008]

Matriella/g [g:2008]  


References. Edmunds 1959 (subgen.): * *; Allen & Edmunds 1965 (subgen.): * *; Edmunds & Jensen & Berner 1976 (subgen.): * *; Kluge 1997d (subgen.): *; Kluge 2004: * * * *


Plesiomorphies of Ephemerella/fg5. Maxilla is biting [see Ephemerella/fg1 (16)], vestige of maxillary palp always present, 3-segmented [see Ephemerella/fg1 (1)]. Larval thorax without projections characteristic for Cincticostella/g1 or Notacanthella. Larval fore legs are non-grasping (unlike Drunella/g1). Larval paracercus has the same length as cerci (unlike Caudatella); secondary swimming setae are present on lateral and medial sides of cerci and on lateral sides of paracercus [see Furcatergaliae (6)] (among Ephemerella/fg3, the same in some Drunella/fg1 and some Torleya/fg1).

Size. Fore wing length 613 mm.

Distribution. Holarctic and Oriental Region.


Nominal species in Ephemerella/fg5:  

alleni Jensen & Edmunds 1966 [Ephemerella]

argo Burks 1949 [Ephemerella] syn.subj. excrucians [Ephemerella]

aronii Eaton (in Esben-Petersen) 1908 [Ephemerella] syn.subj. aurivillii [Chitonophora]

atagosana Imanishi 1937 [Ephemerella] typus nominis Hosoba

aurivillii Bengtsson 1908 [Chitonophora] --/

berneri Allen & Edmunds 1958 [Ephemerella (Chitonophora)] typus nominis Tsalia

catawba Traver 1932 [Ephemerella]

choctawhatchee Berner 1945 [Ephemerella] syn.subj. invaria [Baetis]

cognata Traver 1934 [Ephemerella] syn.subj. teresa [Ephemerella]

concinnata Traver 1934 [Ephemerella] syn.subj. aurivillii [Chitonophora]

crenula Allen & Edmunds 1965 [Ephemerella (Ephemerella)] syn.subj. excrucians [Ephemerella]

dentata Bajkova 1967 [Ephemerella]   syn.subj. atagosana [Ephemerella]

denticula Allen 1971 [Ephemerella (Ephemerella)] syn.subj. atagosana [Ephemerella]

dorothea Needham 1908 [Ephemerella]

euterpe Traver 1934 [Ephemerella] syn.subj. maculata [Ephemerella]

excrucians Walsh 1862 [Ephemerella]

ezoensis Gose 1985 [Ephemerella (Cincticostella)] syn.subj. aurivillii [Chitonophora]

feminina Needham 1924 [Ephemerella] syn.subj. invaria [Baetis]

floripara McCafferty 1985 [Ephemerella] syn.subj. invaria [Baetis]

fratercula McDunnough 1925 [Ephemerella] syn.subj. invaria [Baetis]

hispida Allen & Edmunds 1965 [Ephemerella]

idahoensis Allen 1984 [Caurinella]

inconstans Traver 1932 [Ephemerella] syn.subj. invaria [Baetis]

inermis Eaton 1884 [Ephemerella] syn.subj. excrucians [Ephemerella]

infrequens McDunnough 1924 [Ephemerella]

intermedia Keffermuller 1979 [Ephemerella krieghoffi] syn.subj. mucronata [Chitonophora]

invaria Walker 1853 [Baetis]

keijoensis Allen 1971 [Ephemerella] syn.subj. atagosana [Ephemerella]

kozhovi Bajkova 1967 [Ephemerella  typus nominis Zonadia --/

krieghoffi Ulmer 1920 [Chitonophora] syn.subj. mucronata [Chitonophora]

lacustris Allen & Edmunds 1965 [Ephemerella] syn.subj. excrucians [Ephemerella]

maculata Traver 1934 [Ephemerella]

maxima Allen 1971 [Ephemerella] syn.subj. aurivillii [Chitonophora]

moffatae Allen 1977 [Ephemerella] syn.subj. mucronata [Chitonophora]

mollitia Seemann 1927 [Ephemerella] syn.subj. dorothea [Ephemerella]

mucronata Bengtsson 1909 [Chitonophora] ,/

needhami McDunnough 1925 [Ephemerella] typus nominis Draeconia

norda McDunnough 1924 [Ephemerella] syn.subj. aurivillii [Chitonophora]

notata Eaton 1887 [Ephemerella]

notofascia Yoon & Bae 1988 [Ephemerella] syn.subj. kozhovi [Ephemerella]

ora Burks 1949 [Ephemerella] syn.subj. excrucians [Ephemerella]

rama Allen 1968 [Ephemerella] syn.subj. excrucians [Ephemerella]

rossi Allen & Edmunds 1965 [Ephemerella] syn.subj. excrucians [Ephemerella]

rotunda Morgan 1911 [Ephemerella] syn.subj. invaria [Baetis]

semiflava McDunnough 1926 [Ephemerella] syn.subj. excrucians [Ephemerella]

simila Allen & Edmunds 1965 [Ephemerella] syn.subj. invaria [Baetis]

subvaria McDunnough 1931 [Ephemerella]

taeniata Tshernova 1952 [Ephemerella] syn.subj. aurivillii [Chitonophora]

teresa Traver 1934 [Ephemerella]   typus nominis Matriella

unicolorata Ikonomov 1961 [Chitonophora] syn.subj. mucronata [Chitonophora]

velmae Allen & Edmunds 1963 [Ephemerella (Serratella)]

vernalis Banks 1914 [Ephemerella] syn.subj. invaria [Baetis]

verruca Allen & Edmunds 1965 [Ephemerella]


See also:

Ephemerella/fg3 INCERTAE SEDIS 

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