CLADOENDESIS OF EPHEMEROPTERA

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Tricorythopsis

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Ephemerella/fg1 Pantricorythi Tricoryptera Leptohyphes/fg1 - Tricorythopsis)

Nomen hierarchicum: Tricorythopsis/g(1) [g:1958]

In circumscription fits:

— gen. Tricorythopsis Traver 1958: 491

— Tricorythopsis/g(1): Kluge 2004: 344


References. Traver 1958: ; – Molineri 1999b: ' *; – Wiersema & McCafferty 2000: *; – Molineri 2001c: * *; – Kluge 2004: * *


Autapomorphies of Tricorythopsis.

(1) On fore wing proximal end of iMP is adjacent or connected with CuA, while MP2 originates more distally; thus iMP, MP2 and CuA form a secondary triad (Molineri 1999: Fig.19–37; 2001c: Fig.51–107). In other Pantricorythi MP2 is usually also detached from MP-MP1, but is not shorter than iMP.

(2) On middle and hind legs of imago and subimago tarsus is 4-segmented: it has only 2 articulations between tibio-tarsal articulation [see (7)] and the articulation of claw-bearing segment (initial segment 5) (Molineri 2001c: Fig.20). The same in Tricorygnatha and some others (see Index of characters [2.2.83]).

(3) Gonostyli are non-functional and have peculiar structure: distal segment is lost (non-unique character – see Index of characters [2.3.12]); the last (2nd) segment is directed laterally [see Pantricorythi (2)]; styliger forms a pair of more or less prominent narrow gonostyli pedestals lacking muscles (Molineri 1999b: Fig.18–35; 2001c: Fig.1–102).

(4) Egg [initially having a polar cap – see Ephemerella/fg1 (14)] has 2 polar caps (Molineri 200c: Fig.122–131). Non-unique character (see Index of characters [3.5]).

Characters of Tricorythopsis common with Tricorythodes and Caenoptera.

(5) Lateropostnotum is large, stretches up to sternum and fused with epimeron.

(6) Hind wings [see Leptohyphes/fg1 (2)] are lost in both sexes (see Index of characters [2.2.59]).

(7) On middle and hind legs of imago and subimago [see (2)] 1st tarsal segment [initially fused with tibia and shortened – see Furcatergaliae (2)] on outer side is secondarily distinctly separated from tibia; joining of tibia and tarsus is strongly inclined resembling larval condition, thus outer side of 1st tarsal segment is strongly elongate (Molineri 2001c: Fig.20). The same is Tricorythodes (possibly synapomorphy) and Caenoptera.

Characters of Tricorythopsis of unclear phylogenetic status. 

(8) Tergalial ventral lobes [see Leptohyphes/fg1 (1)] have peculiar structure uniform in all species: on tergalius II (operculate) each of two branches has one process (the same in Leptohyphodes only), costal flap is probably lost; on each tergalius III–IV costal branch (widely projected, as in other Leptohyphes/fg1) has one process arising from base, anal branch is narrow and lacks processes; on tergalius V costal branch is also wide, but lacks process, anal branch is lost; tergalius VI has no ventral lobe (Molineri 2001c: Fig.6–121).

Size. Small, fore wing length 2–3 mm (see Tricoryptera). 

Distribution. Neotropical Region.


Nominal species in Tricorythopsis/g(1):

acara Belmont & Salles & Hamada 2011 [Tricorythopsis]

araponga Dias & Salles 2005 [Tricorythopsis]

artigas Traver 1958 [Tricorythopsis]

bahiensis Dias & Salles & Ferreira 2008 [Tricorythopsis]

baptistai Dias & Salles 2005 [Tricorythopsis]

chiriguano Molineri 2001 [Tricorythopsis]

fictus Molineri 1999 [Tricorythopsis] — syn.subj. artigas [Tricorythopsis]

gibbus Allen 1967 [Leptohyphes]

minimus Allen 1973 [Leptohyphes]

petersorum Molineri 1999 [Tricorythopsis] — syn.subj. undulatus [Leptohyphes]

pseudogibbus Dias & Salles 2005 [Tricorythopsis]

sigillatus Molineri 1999 [Tricorythopsis]

spongicola Lima & Salles & Pinheiro 2011 [Tricorythopsis]

ticuna Molineri & Zuniga 2006 [Tricorythopsis]

undulatus Allen 1967 [Leptohyphes]

viriosus Allen 1973 [Leptohyphes] — syn.subj. minimus [Leptohyphes]

volsellus Molineri 1999 [Tricorythopsis]

yacutinga Molineri 2001 [Tricorythopsis]

yucupe Dias & Salles & Ferreira 2008 [Tricorythopsis]


See also: 

Leptohyphes/fg1 INCERTAE SEDIS

Furcatergaliae INCERTAE SEDIS

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