CLADOENDESIS OF EPHEMEROPTERA

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Cloeodes/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata
 Anteropatellata Baetovectata - Cloeodes/g1)

Nomen hierarchicum: Cloeodes/g1 [g:1938] (incl. ??Bungona, Centoptella)

In circumscription fits:

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References.  


Autapomorphies of Cloeodes/g1.

(1) Larval abdomen and legs with well-developed regular rows of trichobothria (long thin setae in round sockets). Each of abdominal sterna II–VI or some of them can bear a pair of short transverse rows of trichobothria (unique apomorphy); in Cloeodes/g2 such trichobothria are present on all sterna II–VI, while in Centroptella they can be retained on segments IV–VI only (in ...??Bungona) or completely lost (in ...??). Patella-tibial suture bears a row of trichobothria at lest in its proximal part: on posterior side of tibia this row is directed nearly transversely, on anterior side – more longitudinally [see Plesiomorphies of Cloeodes/g2 and Centroptella (1) below]. Tarsus can bear a longitudinal row of trichobothria on outer side. In other Turbanoculata such rows of trichobothria on patella-tibial suture and tarsus can be developed, but usually are not so conspicuous.

Characters of Cloeodes/g1 of unclear phylogenetic status. 

(2) Larval antenna bases are not brought together, frons between them does not form a keel (unlike Paracloeodes and some Baetungulata – see Index of characters [1.1.3]).

() Larval abdominal terga with scales in operculate sockets [see Turbanoculata (12)] (see Index of characters [1.3.4]).

() Tergalii are unable to make rhythmical respiratory movements. Non-unique apomorphy (see Index of characters [1.3.30]).

Plesiomorphies of Cloeodes/g1. Larval paracercus is allways developed.

Variable characters of Cloeodes/g1. 

?Maxillary palp can be 3-segmented or 2-segmented. 

Claws [initially with two rows of denticles – see Tetramerotarsata (11)] can have two rows of prominent or poorly developed denticles (in selected Centroptella), or lack denticles (in Cloeodes/g2 and selected Centroptella). 

On fore wing marginal intercalaries are situated mainly in twos in each field [see Baetovectata (1)]; but sometimes, especially in females, one of two veins is lost, and in each field a single eccentric intercalary is present. 

Hind wings can be present (in Cloeodes/g2 penai [Notobaetis]) or lost (in all other species); in the last case vestiges of hind protoptera can be present or lost. 

In various species, in mature larva ready to moult to subimago, buds of subimaginal gonostyli [see Baetovectata (3)] can be folded under larval cuticle by the "Nigrobaetis-type" (2nd segment directed posteriorly and curved by its convexity medially and by apex laterally) or by the ?? "Baetis-type" (2nd segment straight and directed posteriorly-medially) (see Index of characters [2.3.9]).

Size. Fore wing length 2–10 mm (see Tetramerotarsata).

Distribution. Neotropical, Oriental and Afrotropical Regions.


The taxon Cloeodes/g1 is divided into:

1. plesiomorphon Cloeodes/g2

2. Centroptella/g1