CLADOENDESIS OF EPHEMEROPTERA

ABC

z 

Campsurus/fg3

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Fimbriatotergaliae Fossoriae Cryptoprosternata Polymitarcys/f1=Ephoron/g2  
Campsurus/fg1
AsthenopusCampsurus/fg2 - Campsurus/fg3)

Nomen hierarchicum: Campsurus/fg3 [f:1935; g:1868] (sine Tortopus)

In circumscription fits:

gen. Campsurus: Eaton 1871

Campsurus/fg3: Kluge 2004: 268


References. Lestage 1923b: ; McCafferty 1975: ; Edmunds & Jensen & Berner 1976: * *; Kluge 2004: * * * *


Autapomorphies of Campsurus/fg3.

(1) In male and female imago middle and hind legs [vestigial see Campsurus/g1 (6)] consist of coxa and trochanter only. When larva molts to subimago, its legs distad of trochanter, do not undergo metamortphosis, retain larval musculature up to the ecdysis, and at ecdysis are shed together with remainder exuviae, containing larval muscles inside.

Plesiomorphies of Campsurus/fg3 (unlike Tortopus).  Mandibular tusk [see Campsurus/fg1 (1)] on median margin without subapical denticle; it has either a row of equal small denticles, or a larger denticle near base and a small denticles distad of it. Vestige of tergalius I [see Cryptoprosternata (6)] is bilamellate. Pedestal of gonostylus without pointed lateral process.

Size. Fore wing length 520 mm.

Distribution. America (dominate in Neotropical Region).


Nominal species in Campsurus/fg3: 

 albicans Percheron 1838 [Ephemera]

 albifilum Walker 1853 [Palingenia]

 amapaensis Molineri & Emmerich 2010 [Campsurus]

 argentinus Esben-Petersen 1912 [Campsurus]

 assimilis Traver 1944 [Campsurus]

brasiliensis Traver 1944 [Campsurus] syn.subj. major [Campsurus]

 burmeisteri Ulmer 1942 [Campsurus] ?

 claudus Needham & Murphy 1924 [Campsurus]

 corumbanus Needham & Murphy 1924 [Campsurus]

cuspidatus Eaton 1871 [Campsurus]

 cuyuniensis Traver 1947 [Campsurus]

 dallasi Navás 1927 [Campsurus]

decolorata Hagen 1861 [Palingenia]

 dorsalis Burmeister 1839 [Palingenia]

 duplicatus Spieth 1943 [Campsurus]

emersoni Traver 1947 [Campsurus]

 essequibo Traver 1947 [Campsurus]

 evanidus Needham & Murphy 1924 [Campsurus]

fuliginatus Molineri & Salles 2013 [Campsurus]

gracilipenis Molineri & Salles 2013 [Campsurus]

 holmbergii Weyenbergh 1883 [Ephemera]

homaulos Molineri & Salles 2013 [Campsurus]

 indivisus Ulmer 1942 [Campsurus]

 jorgenseni Esben-Petersen 1912 [Campsurus]

juradinus Navás 1930 [Campsurus] syn.subj. evanidus [Campsurus]

 latipennis Walker 1853 [Palingenia] typus nominis Campsurus

 litaninensis Spieth 1943 [Campsurus]

 longicauda Navás 1931 [Campsurus]

 lucidus Needham & Murphy 1924 [Campsurus]

mahunkai Puthz 1973 [Campsurus] syn.subj. truncatus [Campsurus]

major Needham & Murphy 1924 [Campsurus]

 melanocephalus Pereira & Da-Silva 1991 [Campsurus] syn.subj. truncatus [Campsurus]

meyeri Navás 1934 [Campsurus] syn.subj. violaceus [Campsurus]

molinai Molineri & Salles & Emmerich 2015 [Campsurus]

mutilus Needham & Murphy 1924 [Campsurus]

 nappii Weyenbergh 1883 [Palingenia]

 notatus Needham & Murphy 1924 [Campsurus] syn.subj. violaceus [Campsurus]

 pallidus Needham & Murphy 1924 [Campsurus] syn.subj. argentinus [Campsurus]

paranensis Navás 1933 [Campsurus] syn.subj. violaceus [Campsurus]

 pedicellarius Spieth 1943 [Campsurus]

 pictetii Kirby 1897 [Campsurus]

 pfeifferi Navás 1931 [Campsurus]

 quadridentatus Eaton 1871 [Campsurus]

 scutellaris Needham & Murphy 1924 [Campsurus] syn.subj. jorgenseni [Campsurus]

 segnis Needham & Murphy 1924 [Campsurus]

 striatus Needham & Murphy 1924 [Campsurus]

truncatus Ulmer 1920 [Campsurus]

ulmeri Traver 1950 [Campsurus]

 violaceus Needham & Murphy 1924 [Campsurus]

 vulturorum Emmerich & Molineri 2011 [Campsurus]

wappaei Weyenbergh 1883 [Ephemera]

yavari Molineri & Salles 2013 [Campsurus]

 zikani Navás 1934 [Campsurus]


Examined:

 sp.A (Peru) -;//

 sp.B (Peru)

 sp.C (Peru)

 sp.1 (Peru) exuv.

 sp.2 (Peru) exuv.

 sp.3 (Peru) exuv.

 sp.4 (Peru) exuv.

 sp.5 (Peru) exuv.


See also:

Campsurus/fg1 INCERTAE SEDIS