CLADOENDESIS OF EPHEMEROPTERA

ABC

z 

Tortopus/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Fimbriatotergaliae Fossoriae Cryptoprosternata Polymitarcys/f1=Ephoron/g2  
Campsurus/fg1
AsthenopusCampsurus/fg2 - Tortopus/g1)

Nomen hierarchicum: Tortopus/g1 [g:1924]

In circumscription fits:

gen. Tortopus Needham & Murphy 1924: 23

Tortopus/g(1): Kluge 2004: 268


References. Needham & Murphy 1924: ; Ulmer 1942: ; Traver 1950: ; Burks 1953: ; Scott & Berner & Hirsch 1959: ; Koss 1968: ; McCafferty 1975: ; Edmunds & Jensen & Berner 1976: * *; Dominguez 1987: ; McCafferty & Bloodgood 1989: ; Lugo-Ortiz & McCafferty 1996b: ; Kluge 2004: * * * *


Autapomorphies of Tortopus/g1.

(1) Pedestal of gonostylus [actively movable see Campsurus/fg2 (2)] with a pointed lateral process parastylus (Kluge 2004: Fig.80:CD). Female has receptors of parastyli a pair of impressions on abdominal sternum VIII (McCafferty & Bloodgood 1989: Fig.19). Unique apomorphy.

(2) Mandibular tusk [see Campsurus/fg1 (1)] on its median margin with a subapical denticle, sometimes with one more denticles proximad of it.

(3) Vestige of tergalius I [initially bilamellate see Cryptoprosternata (6)] is unilamellate. Non-unique apomorphy (see Index of characters [1.3.33]).

Plesiomorphy of Tortopus/g1. Unlike Campsurus/fg3, middle and hind legs of male and female imago retain vestigial non-functional femur, tibia and tarsus.

Size. Fore wing length 920 mm.

Distribution. America; dominate in Neotropical Region.


The taxon Tortopus/fg1 is divided into:

1. Tortopus/g2

2. Tortopsis