CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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† Hexagenites/fg1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
INCERTAE SEDIS)
Nomen hierarchicum:
Hexagenites/fg1
[f:1917; g:1880]
In circumscription fits:
— subfam. Ephemeropsinae Cockerell 1924: 136
— fam. Hexagenitidae: Tshernova 1961: 589
— Hexagenites/fg1: Kluge 2004: 353
Synonyms: subjective synonyms of Hexagenites/fg (typus weyenberghii [H.]) with conspecific type-species:
Paedephemera/fg
[f:1917; g:1906] (typus multinervosa [Ephemera])
Stenodicranum/fg
[fg: 1954], typus cellulosa [Ephemera])
(subjective synonymy: cellulosa [E.] = weyenberghii [H.] = multinervosa [E.] (Demoulin 1970c: 7)
Nominal taxa included:
Ephemeropsis/fg
[f:1924; g:1864]
Phacelobranchus/g
[g:1906]
Protoligoneuria/g
[g:1955]
Hexameropsis/g
[g:1974]
Libanoephemera/g
[g:2019]
Mongologenites/g
[g:1986]
Palaeobaetodes/g
[g:1987]
Cratogenites/g
[g:1996]
Cratogenitoides/g
[g:1996]
Cratohexagenites/g [g:2007]
Caenoephemera/g
[g:2001]
Baikalogenites/g [g:2017]
?
Shantous/g
[g:2007] 
Epicharmeropsis/g [2007]
Possibly, here also belongs Costalimella (see Euplectoptera incertae sedis).
References.
Cockerell 1924: 
;
– Tshernova 1961: *;
Meshkova 1961: ; –
Tshernova 1970: ;
– Tshernova & Sinitshenkova 1974: ';
– McCafferty 1990: ';
– Martins-Neto 1996: ';
– Kluge 2004: * *
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Autapomorphies of Hexagenites/fg1. (1) On fore wing CuA bifurcates to CuA1 and CuA2 (non-unique character – see Index of characters [2.2.50]), from the bifurcation a vein iCu arises, which forms several (3–5) triads following one another: anterior branch of each triad forms next triad; each of these triads has anterior branch arched by its convexity anteriorly; all branches of these triads go to basitornal margin of wing. Unique apomorphy. (2) Fore and hind wings with more or less expressed vein gemination: at least there are pairs of veins brought together: RSp+MA1, iMA+MA2, MP2+CuA; MP1 and iMP are more or less approximated to iMA+MA2. Non-unique apomorphy (see Index of characters [2.2.28]). (3) Tergalius VII is enlarged, distinctly longer than others, while tergalii I-VI have subequal length (unique apomorphy). On all tergalii at least costal rib is thickened and anal rib is situated at a distance from anal margin (as in some other mayflies – see Index of characters [1.3.28]). Plesiomorphies of Hexagenites/fg1. Larva has siphlonuroid appearance, with dense primary swimming setae on caudalii. |
Size. Fore wing length 16–42 mm.
Age. Late Jurassic – Late Cretaceous (Europe, Asia, South America).
Nominal species in Hexagenites/fg1:
firmus Sinitshenkova 2017 [Baikalogenites]
— typus nominis Baikalogenites
berkeyi
Cockerell 1927 [Ephemeropsis trisetalis]
braueri Handlirsch 1906 [Phacelobranchus]
— syn.subj. trisetalis [Ephemeropsis]
branchicillus Huang & Sinitshenkova
& Ren 2011 [Siberiogenites]
britoi Martins-Neto 1996 [Palaeobaetodes]
cellulosa Hagen 1862 [Ephemera]
— typus nominis Stenodicranum
chaneyi
Cockerell 1927 [Ephemeropsis trisetalis]
corradiniae
Martins-Neto 1996 [Cratogenites]
costalimai Brito 1987 [Palaeobaetodes]
delclosi Martins-Neto 1996 [Cratogenitoides]
elongatus Lin
& Kong & Shih & Zhao
& Ren 2018 [Hexameropsis]
heloisae
Storari & Godunko
& Salles & Saraiva & Staniczek & Rodrigues 2021 [Protoligoneuria]
hexavenulosus
Huang & Ren & Sinitshenkova
& Shih 2007 [Epicharmeropsis]
— typus nominis Epicharmeropsis
inopinatabranchia Azar
& Ferté & El Hajj & Nal & Maksound 2019 [ Libanoephemera]
— typus nominis Libanoephemera
khasurtensis Sinitshenkova 2017 [Furvoneta]
lacustris Zhang & Kluge 2007 [Shantous] —
typus nominis Shantous
lagueatus Sinitshenkova 1986 [Mongologenites]
limai Demoulin 1955 [Protoligoneuria]
longicercus
Staniczek 2007 [Cratohexagenites]
— typus nominis Cratohexagenites
martynovae
Tshernova 1961 [Ephemeropsis]
melanurus
Cockerell 1924 [Ephemeropsis]
middendorffii
Jones [Estheria]
middendorfi Handlirsch 1906 [Ephemeropsis]
— syn.subj. trisetalis [Ephemeropsis]
minor
Staniczek 2007 [Cratohexagenites]
mortua
Hagen 1862 [Ephemera]
— syn.subj. cellulosa [Ephemera]
multinervosa Oppenheim 1888 [Ephemera]
— typus nominis Paedephemera
— syn.subj. cellulosa [Ephemera]
oppenheimi
Handlirsch 1906 [Paedephemera]
— syn.subj. cellulosa [Ephemera]
orientalis
Eichwald 1868 [Ephemeropsis]
— syn.subj. trisetalis [Ephemeropsis]
quadrivenulosus
Huang & Ren & Sinitshenkova
& Shih 2007 [Epicharmeropsis]
reisi Handlirsch 1939 [Ephemeropsis]
— syn.subj. trisetalis [Ephemeropsis]
rossica Eichwald 1866 [Lethaea]
— syn.subj. trisetalis [Ephemeropsis]
selini Tshernova &
Sinitshenkova 1974 [Hexameropsis]
shangyuanensis Lin & Huang
2001 [Caenoephemera]
trisetalis Eichwald 1864 [Ephemeropsis]
weyenberghii Scudder 1880 [Hexagenites]
— syn.subj. cellulosa [Ephemera]
See also:
other Euplectoptera
INCERTAE SEDIS
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