Heptagenia

CLADOENDESIS OF EPHEMEROPTERA
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Heptagenia/f8=g7

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata
Heptagenia/f5=g4_{Ecdyonurus/fg1}Heptagenia/f6=g5_KageroniaHeptagenia/f7=g6_{Stenonema/fg1}Heptagenia/f8=g7)

Nomen hierarchicum: Heptagenia/f8=g7 [f:1901; g:1863] (sine Dacnogenia, Stenonema; incl. Sigmoneuria, Trichogenia)

In circumscription fits:

— subgen. Heptagenia: Kluge 1988: 295

— Heptagenia/f8=g7: Kluge 2004: 169

Nominal taxa included:

Sigmoneuria/g [g:1964]

Trichogenia/fg [f:2006; g:1988]

References. Kluge 1988: *; – 1993: *; – 2004: * * * *.

Characters of Heptagenia/f8=g7 of unclear phylogenetic status.

(1) Superlingua [curved laterally – see Heptagenia/f5=g4 (1)] on its lateral margin with a subapical incision and a convexity proximad of it (Kluge 2004: Fig.55:F); by its form differs from any other taxon among Heptagenia/f5=g4.

(2) Each glossa [initially pyramidal – see Radulapalpata (1)] is narrow, curved, in its distal half flattened, as dorsal ridge is approximated to median edge (Kluge 2004: Fig.55:D-E). The same in Stenonema/fg1; probably synapomorphy.

(3) In imago and subimago 1^st tarsal segment is shortened, on fore tarsus of male several times shorter than 2^nd segment.

Non-unique apomorphy (see Index of characters [2.2.76]); among Pentamerotarsata the same in Dacnogenia and Raptoheptagenia, in selected species of Kageronia and Stenonema/fg1, in some Ecdyonurus/fg1 and Rhithrogena/fg3.

Plesiomorphy of Heptagenia/f8=g7. Unlike Dacnogenia, hypopharynx is not shortened, has the same length as superlinguae [see (1)] (Kluge 2004: Fig.55:F).

Variable characters of Heptagenia/f8=g7. Tergalii [see Branchitergaliae (3)] are usually non-modified: usually all tergalii I–VII with fibrillose lobe (only on maxillaris [T.] tergalius VII lacks fibrillose lobe); anal and costal ribs are either located on anal and costal margins, or lost.

Size. Fore wing length 7–15 mm.

Distribution. Holarctic and Oriental Region.

Nominal species in Heptagenia/f8=g7:

adaequata McDunnough 1924 [Heptagenia]

albicauda Kluge 1987 [Heptagenia sulphurea] --/

amseli Demoulin 1964 [Sigmoneuria] — typus nominis Sigmoneuria — syn.subj. perflava [Heptagenia]

arsenjevi Tshernova 1952 [Heptagenia] — syn.subj. flava [Heptagenia]

bipunctata Esben-Petersen 1916 [Heptagenia] — syn.subj. flava [Heptagenia]

caudata [Baetis] nonen nudum — syn.obj. longicauda [Baetis]

cerea Pictet 1843 [Baetis] — syn.subj. longicauda [Baetis]

chinensis Ulmer 1920 [Heptagenia] --/

citrina Hummel 1825 [Ephemera] nomen oblitum — syn.subj. flava [Heptagenia]

costalis Curtis 1834 [Baetis] — syn.subj. sulphurea [Ephemera]

couzonica Real 1947 [Heptagenia sulphurea]

coxalis Banks 1914 [Heptagenia] — syn.subj. elegantula [Rhithrogena]

culacantha Evans & Botts & Flowers 1985 [Heptagenia]

cyanops Pictet 1843 [Baetis] — syn.subj. sulphurea [Ephemera]

dalecarlica Bengtsson 1912 [Heptagenia] — forma in sulphurea [Ephemera] --/

denervosa Tiensuu 1935 [Heptagenia sulphurea]

diabasia Burks 1947 [Heptagenia] — syn.subj. elegantula [Rhithrogena]

dolosa Traver (in Needham & Traver & Hsu) 1935 [Heptagenia]

elegans Curtis 1834 [Baetis] — syn.subj. sulphurea [Ephemera]

elegantula Eaton 1885 [Rhithrogena]

ferruginea Gmelin 1790 [Ephemera] — syn.subj. sulphurea [Ephemera]

flava Rostock 1878 [Heptagenia] --/

flavata Navás 1922 [Heptagenia]

flavescens Walsh 1862 [Palingenia] — typus nominis Heptagenia

flavipennis Dufour 1841 [Ephemera] — syn.subj. longicauda [Baetis]

grandis [Ecdyonurus] nomen nudum — syn. pullus [Ecdyonurus]

guranica Belov 1981 [Heptagenia] --/

helvola Sulzer 1776 [Ephemera] — syn.subj. sulphurea [Ephemera]

hubleyi Webb & McCafferty (in Webb & Braasch & McCafferty) 2006 [Trichogenia]

julia Traver 1933 [Heptagenia]

kyotoensis Gose 1963 [Heptagenia]

leucophthalma Ström 1783 [Ephemera] — syn.subj. sulphurea [Ephemera]

longicauda Stephens 1835 [Baetis] --/

lutea Kluge 1987 [Heptagenia (Heptagenia)] nom. praeocc. — syn.subj. samochai [Sigmoneuria]

marginalis Banks 1910 [Heptagenia]

maxillaris Braasch & Soldán 1988 [Trichogenia]

patoka Burks 1947 [Heptagenia]

perflava Brodsky 1930 [Heptagenia] --/

platysma Tshernova 1941 [Heptagenia] — syn.subj. perflava [Heptagenia]

pullus Clemens 1913 [Ecdyurus]

quadripunctata Kluge 1989 [Heptagenia (Heptagenia)] ,--

querula McDunnough 1924 [Heptagenia] — syn.subj. elegantula [Rhithrogena]

samochai Demoulin 1973 [Sigmoneuria] --/

soldatovi Tshernova 1952 [Heptagenia] — syn.subj. sulphurea [Ephemera]

solitaria McDunnough 1924 [Heptagenia]

straminea Curtis 1834 [Baetis] — syn.subj. sulphurea [Ephemera]

sulphurea Müller 1776 [Ephemera] --/

tadzhikorum Tshernova 1931 [Heptagenia] — syn.subj. perflava [Heptagenia]

townesi Traver (in Needham & Traver & Hsu) 1935 [Heptagenia]

traverae Braasch 1986 [Heptagenia]

ulmeri Brasch & Webb (in Webb & Braasch & McCafferty) 2006 [Trichogenia]

whitingi Webb & McCafferty (in Webb & Sun & McCafferty & Ferris) 2007 [Heptagenia]