CLADOENDESIS OF EPHEMEROPTERA

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Austremerella

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Ephemerella/fg1 Pantricorythi - Austremerella)

Nomen hierarchicum: Austremerella/fg(1) [f:1997; g:1963]

In circumscription fits:

— subgen. Austremerella: Allen 1965: 264

— gen. Austremerella Riek 1963: 50

— Austremerella/fg(1): Kluge 2004: 317


References. Riek 1963: ; – Allen 1965: ; – Riek 1970:  ; – Peters & Campbell 1991: ; – McCafferty & Wang 1997: ; – Kluge 2004: ; – Suter & Mynott 2013:


Characters of Austremerella of unclear phylogenetic status. 

(1) Tergalii I [initially stick-like – see Ephemerella/fg1 (13)] are lost. Non-unique apomorphy (see Index of characters [1.3.33]).

(2) Tergalius VI has ventral lobe non-bifurcate, thus only tergalii II–V have bifurcate ventral lobes [see Ephemerella/fg1 (7)] (Kluge 2004: Fig.95:F–H). The same structure of tergalius VI occurs in Ephemerella/fg4 only. Possibly, synapomorphy with Melanemerella/f1=Teloganodes/g1 and/or Tricoryptera, where ventral lobe of tergalius VI is vestigial or lost (see Table and Index of characters [1.3.57]).

(3) Tergalii VII [small, with non-bifurcate ventral lobes – see Ephemerella/fg1 (7)] are strongly shifted anteriorly, being located at midlength of tergite (Kluge 2004: Fig.95:H) (unlike Vietnamella and Ephemerella/fg2, whose tergalii VII retain position close to posterior margin of tergite). In Tricoryptera and Melanemerella/f1=Teloganodes/g1 these tergalii are lost, thus their initial position is unknown.

(4) Fore wing with free marginal intercalaries (as in many other Ephemerella/fg1 – see Index of characters [2.2.55]).

Plesiomorphies of Austremerella. Larval head, mandibles and fore legs without modifications peculiar for Vietnamella. Larval abdominal terga with paired submedian projections [see Ephemerella/fg1 (19)] (Riek 1963: Fig.1). Unlike Melanemerella/f1=Teloganodes/g1 and some others, tergalii II are not operculate. Cubital field of fore wing has up to 2 bifurcate veins arising from CuA [see Ephemerella/fg1 (15)]; hind wing is well-developed, as long as 0.2 of fore wing length, with Sc reaching wing apex, RS branching, and veins behind MP1 developed (Riek 1963: Fig.10) (unlike Teloganodes/f1=g3, Teloganella and Tricoryptera).

Size. Fore wing length 12 mm.

Distribution. Australia (this is the only representative of Ephemerella/fg1 in Australian Region).


Nominal species in Austremerella/g(1):

picta Riek 1963 [Austremerella]


See also: 

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