CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Plesiomorphon Labiobaetis/f2=g1
(Panephemeroptera
Euephemeroptera
Euplectoptera Anteritorna
pm.Tridentiseta
Tetramerotarsata
Liberevenata
Turbanoculata
Anteropatellata Baetovectata
Baetungulata
Labiobaetis/f1=Pseudopannota/g1 -
Labiobaetis/f2=g1)
Nomen hierarchicum:
Labiobaetis/f2=g1
[f:2016; g:1987(No.4)] (sine Pseudopannota)
In circumscription fits:
— gen. Labiobaetis Novikova & Kluge 1987: 13
— Labiobaetis/g1: Kluge 2009:109-134: 117
Nominal taxa included:
Müllerbaetis/g
[g:1994]
Cymulabaetis/g(1)
[g:1995]
References.
Kluge & Novikova 2016: 
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Plesiomorphies and variable characters of Labiobaetis/f2=g1. (1) In larval labium, paraglossae are more or less strengthened and widened, and paraglossae are diminished (Kluge & Novikova 2016: Fig.126). The same in some other taxa. (2) Labial palp has large apical-medial projection (Kluge & Novikova 2016: Fig.126). The same in some other taxa. (3) Prostheca of right (as well as left) mandible always stick-shape, denticulate apically [see Turbanoculata (6)] (unlike Takobia/g1 et al. – see Index of characters [1.1.25]); inner margin of both mandibles between prostheca and mola always without setae (unlike Cloeon/fg1 et al. – see Index of characters [1.1.23]). (4) Larval antennae bases are brought together in variable degree, frons between them elevated and forms more or less expressed keel. The same in some other taxa (see Index of charactes [1.1.3]). (5) Hind wing [see Turbanoculata (10)] if present, is of "Acentrella-type": not wide (narrower than in Baetofemorata, but wider than in Acentrella/fg1), only with 2 longitudinal veins without furcation; costal projection is usually absent, only calcaratus [Baetis] has small costal projection. In many species hind wings are lost (see Index of characters [2.2.59-60]). (6) In larva, on inner side of femur near base, at the place where Baetofemorata has a field of small setae pressed together [comp.: Baetofemorata (1)], small setae can be completely absent, or several separate setae are present, or there is a field of setae brought together; in the last case these setae are not pressed together (there can be only groups of 2–4 setae pressed together), unlike Baetofemorata. All these variants can be found in different specimens or on different legs of the same species (particularly, in atrebatinus [B.]). (7) Larval paracercus is developed. (8) In larva, segments of caudalii can be uniform, or (in desertus [Baetis]) each 4th or each 2nd segment differs by darkened and large denticles (the same in Cloeon/fg1 and some other groups). |
Size. Fore wing length 2–6 mm (see Tetramerotarsata).
Distribution. Afrotropical, Oriental, Holarctic Regions and Australia.
Nominal species in Labiobaetis/f2=g1:
academicus Kaltenbach & Surbakti & Kluge
(in Kaltenbach & Surbakti & Kluge & Gattolliat J& Sartori & Balke)
2021
[Labiobaetis] 
-
-
/
acei
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
alahmadii Gattolliat & Al Dhafer (in Gattolliat & Kondratieff
& Kaltenbach & Al Dhafer) 2018 [Labiobaetis]
aldabae
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
altus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
amber Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
ancoralis Shi
& Tong 2014 [Labiobaetis]
apache McCafferty & Waltz 1995 [Labiobaetis]
aquacidus Lugo-Ortiz & McCafferty 1997
[Labiobaetis]
— syn.subj. latus [Baetis]
arfak Kaltenbach & Gattolliat 2021
[Labiobaetis]
atrebatinus Eaton 1870 [Baetis]
— typus
nominis Labiobaetis
--/
australis Traver 1932 [Baetis]
— syn.subj. frondalis
[Baetis]
baeticatus Burks 1953 [Baetis]
— syn.subj. frondalis
[Baetis]
baganii
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
bakerae Kaltenbach &
Gattolliat 2020
[Labiobaetis]
balcanicus Müller-Liebenau & Soldan 1981 [Baetis]
— typus
nominis Cymulabaetis
balkei Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
batakorum Kaltenbach &
Gattolliat 2019
[Labiobaetis]
bellus Barnard 1932 [Baetis] --/
bilibil Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
boettgeri Ulmer 1924 [Pseudocloeon]
borneoensis Müller-Liebenau 1984 [Baetis]
boussoulius Gillies 1993 [Baetis]
-
-/
branchiaesetis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
brao Kaltenbach & Garces
& Gattolliat 2022
[Labiobaetis]
calcaratus Keffermuller 1972 [Baetis] --/
camiguinensis
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
catadupa Kaltenbach & Gattolliat 2021
[Labiobaetis]
centralensis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
claudiae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
cleopatrae Thomas & Soldan 1989 [Baetis]
dambrensis
Gattolliat 2001 [Labiobaetis]
dardanus McDunnough 1923 [Baetis]
davamanii Sivaruban & Srinivasan
& Barathy & Isack 2022
[Labiobaetis]
dayakorum Kaltenbach &
Gattolliat 2020
[Labiobaetis]
dalisay
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
delocadoi
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
dendrisetis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
desertus Novikova & Kluge 1987 [Baetis (Labiobaetis)]
--/
difficilis Müller-Liebenau 1984 [Baetis]
diffundus Müller-Liebenau 1984 [Baetis]
dipsicus Gillies 1949 [Baetis]
ediai Kaltenbach & Gattolliat 2021 [Labiobaetis]
elachistus Burks 1953 [Baetis]
— syn.subj. dardanus
[Baetis]
elisae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
elouardi Gillies 1993 [Baetis]
ephippiatus Traver (in Needham & Traver & Hsu) 1935 [Baetis]
excavatus Kaltenbach
& Gattolliat 2021 [Labiobaetis]
fabulosus Lugo-Ortiz & McCafferty 1997
[Labiobaetis] 
fastigatum Kopelke 1980 [Pseudocloeon]
freitagi
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
frondalis McDunnough 1925 [Baetis]
fulmeki Ulmer 1939 [Acentrella]
gamay
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
gambiae
Gillies 1993 [Baetis]
geminatus Müller-Liebenau & Hubbard 1985
[Baetis] 
-
-
/
gilliesi
Gattolliat 2001 [Labiobaetis]
gindroi Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
giselae
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
gladius Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
glaucus Agnew 1961 [Baetis] 
grandiculum Kopelke 1980 [Pseudocloeon]
gueuningi Kaltenbach &
Gattolliat 2019
[Labiobaetis]
hattam Kaltenbach & Gattolliat 2021
[Labiobaetis]
hypodeum Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
inconspicuum Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
inopinatus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
inopinum Gillies 1949 [Pseudocloeon]
insolitus Kopelke 1980 [Baetis]
involutum Lugo-Ortiz & McCafferty 1999 (in Lugo-Ortiz &
McCafferty & Waltz) [Pseudocloeon]
itineris Kaltenbach &
Gattolliat 2019
[Labiobaetis]
jacobusi Kubendran & Balasubramanian (in Kubendran &
Balasubramanian & Selvakumar & Gattolliat & Sivaramakrishnan) 2015 [Labiobaetis]
janae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
jonasi Kaltenbach &
Gattolliat 2019
[Labiobaetis]
kalengoensis Kopelke 1980 [Baetis]
kinibeli Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
kokoda Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
kui Kaltenbach & Garces
& Gattolliat 2022
[Labiobaetis]
lachicae
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
latus Agnew 1961 [Baetis] --/
lobatus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
longicercus
Gattolliat 2001 [Labiobaetis]
longipalpus Morihara & McCafferty 1979 [Baetis]
lubu Kaltenbach &
Gattolliat 2019
[Labiobaetis]
macrospinosus Koch 1985 [Baetis]
magnovaldus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
masai Lugo-Ortiz & McCafferty 1997
[Labiobaetis]
mendozai
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
minang Kaltenbach &
Gattolliat 2019
[Labiobaetis]
michaeli
Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
molawinensis Müller-Liebenau 1982 [Baetis]
— typus
nominis Müllerbaetis
mongolicus Tshernova 1952 [Baetis] — syn.subj.
tricolor [Baetis]
moriharai Müller-Liebenau 1984 [Baetis]
morus Chang & Yang (in Kang & Chang & Yang) 1994 [Baetis (Mullerbaetis)]
— syn.subj.
atrebatinus [Baetis]
mtonis Gillies 1994 [Baetis] --/
multus Müller-Liebenau 1984 [Baetis]
mustus Kang & Yang 1996 [Baetis (Labiobaetis)]
nabire Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
nadineae Lugo-Ortiz & McCafferty 1997
[Labiobaetis]
necopinatus Müller-Liebenau 1981 [Baetis]
neglectus Navas 1913 [Baetis]
nigrocercus
Gattolliat
2001 [Labiobaetis]
numeratus Müller-Liebenau 1984 [Baetis]
obscuriventris Tshernova 1952 [Baetis]
— syn.subj. tricolor [Baetis]
onim Kaltenbach & Gattolliat 2021
[Labiobaetis]
operosus Müller-Liebenau 1984 [Baetis]
ordinatus Müller-Liebenau & Hubbard 1985 [Baetis] --
orientalis
Kluge 1983 [Baetis atrebatinus]
— syn.subj.
atrebatinus [Baetis]
orientis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
pakpak Kaltenbach &
Gattolliat 2019
[Labiobaetis]
palawano
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
palmyrae Gillies 1949 [Baetis]
pangantihoni
Kaltenbach & Garces & Gattolliat 2020 [Labiobaetis]
papuaensis
Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
paradiffundus Kaltenbach &
Gattolliat 2019
[Labiobaetis]
paradisus Gattolliat & Staniczek 2011 [Labiobaetis]
paranumeratus Kaltenbach &
Gattolliat 2019
[Labiobaetis]
paraoperosus Kaltenbach &
Gattolliat 2019
[Labiobaetis]
paravitilis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
paravultuosus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
pelingeni
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
penan Kaltenbach &
Gattolliat 2020
[Labiobaetis]
petersorum Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
pilosus Kaltenbach &
Gattolliat 2019
[Labiobaetis]
pindaundensis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
piscis Lugo-Ortiz & McCafferty 1997
[Labiobaetis]
planus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
plectile Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
plumbago
Lugo-Ortiz & McCafferty 1997 [Labiobaetis]
podolakae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
potamoticus Gattolliat & Al Dhafer (in Gattolliat & Kondratieff
& Kaltenbach & Al Dhafer) 2018 [Labiobaetis]
propinqua Walsh 1863 [Cloe] --/
pulchellus Müller-Liebenau & Hubbard 1985 [Baetis] --/
punctatus Gattolliat
2001 [Labiobaetis]
quintus Agnew 1961 [Baetis]
rimba Kaltenbach &
Gattolliat 2019
[Labiobaetis]
roulade Kaltenbach &
Gattolliat 2019
[Labiobaetis]
rutschmannae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
sabordoi
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
schwanderae Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
seramensis Kaltenbach &
Gattolliat 2019
[Labiobaetis]
simbuensis Kaltenbach & Vuataz & Gattolliat 2023
[Labiobaetis]
soldani Kubendran & Rathinakumar & Balasubramanian &
Selvakumar & Sivaramakrishnan 2014 [Labiobaetis]
— syn.subj. pulchellus [Baetis]
sonajuventus Webb 2013 [Labiobaetis]
spatulatus Gillies 1994 [Baetis]
— syn.subj. vinosum [Pseudocloeon]
spinosus McDunnough 1925 [Baetis]
— syn.subj. propingua
[Cloe]
stagnum Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
sulawesiensis Kaltenbach &
Gattolliat 2019 [Labiobaetis]
sumbensis Kaltenbach &
Gattolliat 2019
[Labiobaetis]
sumigarensis Müller-Liebenau 1982 [Baetis]
tagbanwa
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
tenuicrinitum Kopelke 1980 [Pseudocloeon]
— syn.subj. vinosum [Pseudocloeon]
toraja Kaltenbach & Gattolliat 2021
[Labiobaetis]
tricolor Tshernova 1928 [Baetis] --/
tripunctatus Gillies 1994 [Baetis] --/
tuberpalpus Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
ulmeri Müller-Liebenau 1981 [Baetis]
valdezorum
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
vallus Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
vinosum Barnard 1932 [Pseudocloeon] --/
vitile Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
vulgaris Gattolliat
2001 [Labiobaetis]
vultuosum Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon] /
wahai Kaltenbach &
Gattolliat 2019
[Labiobaetis]
wantzeni
Kaltenbach & Garces
& Gattolliat 2020
[Labiobaetis]
weifangae Kaltenbach &
Gattolliat 2019
[Labiobaetis]
werneri Kaltenbach & Gattolliat 2021
[Labiobaetis]
--/
wilhelmensis Kaltenbach &
Gattolliat 2018
[
Labiobaetis]
xeniolum Lugo-Ortiz & McCafferty (in Lugo-Ortiz &
McCafferty & Waltz) 1999 [Pseudocloeon]
Examined also:
sp. "long-Rwenzori" (Uganda) --/
sp. "long-Elgon" (Uganda) --/
sp. "4 blanks" (cf. gambiae) (Uganda)
--/
sp. "Elgon" (Uganda) --/
sp. "toadstool" (Uganda) --/
sp. "white back" (cf. masai) (Uganda)
--/
sp. "Bujagali" (Uganda) --/
spp. (Uganda) ,
sp. (Java) --/
sp. (Sulawesi: Mamasa) --/
sp. "alibaba" (Sulawesi: Pinrang) --/
sp. "VIII-IX" (Sulawesi: Pinrang) --/
spp. --/
See also:
Baetovectata
INCERTAE SEDIS 
Turbanoculata
INCERTAE SEDIS
