CLADOENDESIS OF EPHEMEROPTERA

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Baetofemorata, or Baetis/fg8

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata  
Anteropatellata Baetovectata Baetungulata - Baetofemorata)

Nomen circumscribens: Baetofemorata Kluge & Novikova 2011

Nomen hierarchicum: Baetis/fg8 [fg:1815] (sine Labiobaetis, Nigrobaetis, Camelobaetidius, Baetodes, Fallceon, Caribaetis, Americabaetis; incl. Acentrella, Patites)

In circumscription fits:

— grex subgenn. Baetis: Novikova & Kluge 1994: 624

— Baetofemorata = Baetis/fg8: Kluge & Novikova 2011


References. Waltz & McCafferty 1987:667-670: *; – Waltz & McCafferty 1987:553-568: *; – Novikova & Kluge 1994: * *; – Gaino & Rebora 1999: *; – Kluge & Novikova 2011: *, *


Autapomorphies of Baetofemorata.

(1) In larva, each femur in its proximal part of inner side bears a villopore – a patch of small (0.02 mm long) bristles closely pressed together (Waltz & McCafferty 1987:553-568: Figs.1,4,5). Unique apomorphy. Besides Baetofemorata, similar bristles occur in some species of Labiobaetis/f2=g1, being less closely accumulated.

Characters of Baetofemorata of unclear phylogenetic status.

(2) On all legs of subimago each tarsal segment is covered mainly by blunt microlepides; only in distal part of each segment pointed microlepides can be present [as on segments 1–4 of male fore tarsus – see Baetungulata (4)]. The same in some other taxa (see Index of characters [2.2.83]).

(3) In mature larva ready to moult to subimago, buds of subimaginal gonostyli are folded under larval cuticle [see Turbanoculata (13) and Baetovectata (3)] by the "Baetis-type" or "Acentrella-type": in the both cases both 2nd and 3rd segments of gonostylus are directed medially. In most other Turbanoculata the gonostyli buds are directed more caudally or laterally (see Index of characters [2.3.9]).

(4) Larval antennae bases never brought together, frons between them does not form a keel (see Index of characters [1.1.3]).

(5) Prostheca of right (as well as left) mandible is always stick-shape, denticulate apically [see Turbanoculata (6)] (unlike Takobia/g1 et al. – see Index of characters [1.1.25])

(6) Inner margin of both mandibles between prostheca and mola always without setae (unlike Cloeon/fg1 et al – see Index of characters [1.1.23]).

(7) Labial palp, being 3-segmented, usually has 2nd segment widened or with inner-apical projection [see Index of characters [1.1.58]).

Size. Fore wing length 3–11 mm (see Tetramerotarsata).

Distribution. Arctogea (Holarctic, Oriental and Afrotropical Regions).


The taxon Baetofemorata (or Baetis/fg8) is divided into:

1. Baetis/fg9  

2. Rhodobaetis/g(1)

3. Glossidion/g(1)

4. Tenuibaetis/g(1)

5. Baetiella/g1

6. Patites/g(1)

7. Plauditus/g(1)

8. Acentrella/g1

? Heterocloeon/g1


Baetofemorata incertae sedis