CLADOENDESIS OF EPHEMEROPTERA

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Kageronia

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata
Rhithrogena/fg1Heptagenia/f5=g4Ecdyonurus/fg1Heptagenia/f6=g5 - Kageronia)

Nomen hierarchicum: Kageronia/fg(1) [f:2004; g:1931]

In circumscription fits:

— fuscogrisea-Gruppe: Bogoescu & Tabacaru 1962: 280

— subgen. Parastenacron (manuscript name): Kluge 1983b:16

— subgen. Kageronia: Kluge 1988: 293

— gen. Kageronia: Jacob & Dorn & Haybach 1996: 96

— Kageronia/fg(1): Kluge 2004: 168

Nominal taxon included: Parastenacron – unavailable name


References. Kluge 1988: *; – 1993: *; – 2004: * * * *.


Characters of Kageronia of unclear phylogenetic status.

(1) Superlingua [curved laterally – see Heptagenia/f5=g4 (1)] with slightly concave lateral margin and rounded apex (Kluge 2004: Fig.54:B); by its form differs from any other taxon among Heptagenia/f5=g4.

(2) Glossa [see Radulapalpata (1)] with straight median margin and roundish-convex lateral margin, dorsal edge is not approximated to median edge (Kluge 2004: Fig.54:C,K-L) (unlike Heptagenia/f8=g7, Stenonema/fg1 and Ecdyonurus/fg1).

(3) 2nd+3rd segment of labial palp [specialized as scraping – see Radulapalpata (1)] is pointed, thus the field of scraping setae terminates at a distance from the end of the sclerotized ridge (Kluge 2004: Fig.54:D). The same in Ecdyonurus/fg1 and Rhithrogena/fg1; probably symplesiomorphy. Heptagenia/f7=g6 and Cinygma have different shape of this segment.

(4) Larval femur on its outer margin has no long hairs and long stout setae; only short stout spine-like setae and short hairs are present (Kluge 2004: Fig.54:H) [see Plesiomorphies of Heptagenia/f6=g5]. Probably plesiomorphy.

(5) Larval claws with inner margin denticulate (Kluge 2004: Fig.54:G). Probably plesiomorphy [see Pentamerotarsata (4)] (unlike all other Radulapalpata). Subapical denticles on anterior (dorsal) side of claw are also present (that is usual for many Pentamerotarsata).

(6) Tergalius VII lacks fibrillose ventral lobe, thus the fibrillose ventral lobe [see Branchitergaliae (3)] is present on tergalii I–VI only. Non-unique apomorphy (see Index of characters [1.3.59]); among Heptagenia/f5=g4 the same in Stenonema/fg1 and majority of Ecdyonurus/fg1.

(7) Tergalii are pointed apically (Kluge 1987a: Fig.8–11,17). Non-unique character.

Plesiomorphy of Kageronia. On tergalii costal and anal ribs are located on costal and anal margins correspondingly [see Branchitergaliae (3)] (unlike Stenonema/fg1).

Variable characters of Kageronia. On larval caudalii primary swimming setae [vestigial – see Pentamerotarsata (6)] can be present (in kihada [Heptagenia] and orbiticola [H.]) or lost (in fuscogrisea [Ephemera]). In imago and subimago on fore tarsus 1st segment can be shortened (like in Heptagenia/f8=g7) (in kihada [H.] and orbiticola [H.]) or not shortened (in fuscogrisea [E.]).

Size. Fore wing length 7–16 mm.

Age and distribution. Palaeogene – recent; the recent species fuscogrisea [E.] is known from Baltic amber (Kluge 1986). Palaearctic; all three species occur in Eastern Palaearctic.


Nominal species in Kageronia/fg(1): 

confinis Tshernova 1928 [Ecdyonurus] — syn.subj. fuscogrisea [Ephemera]

convergens Lestage 1924 [Ecdyonurus] — syn.subj. fuscogrisea [Ephemera]

fuscogrisea Retzius 1783 [Ephemera] --/

kihada Matsumura 1931 [Heptagenia] ,

orbiticola Kluge 1987 [Heptagenia (Kageronia)] --/

rossicus Tshernova 1928 [Ecdyonurus] — syn.subj. fuscogrisea [Ephemera]

suzukiella Matsumura 1931 [Kageronia] — typus nominis Kageronia — syn.subj. kihada [Heptagenia]

volitans Eaton 1870 [Heptagenia] — syn.subj. fuscogrisea [Ephemera]


See also: 

Radulapalpata INCERTAE SEDIS  

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