CLADOENDESIS OF EPHEMEROPTERA

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Behningia/fg4

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Fimbriatotergaliae Fossoriae  
Behningia/fg1
ProtobehningiaBehningia/fg2DolaniaBehningia/fg3 ParadolaniaBehningia/fg4)

Nomen hierarchicum: Behningia/fg4 [f:1937; g:1930] (sine Paradolania)

In circumscription fits:

— gen. Behningia Lestage 1930: 436, s.str. (sensu Zheng & Gong & Qiang & Zhou 2024)

— Behningia/fg3: Kluge 2004

— Behningia/fg4


References. Ulmer 1924d ("Ephemeriden-Nymphe"): ; – Motas & Bacesco 1937: ; – Tshernova 1938: ; – 1952: ; – Keffermuller 1959: ; – Edmunds & Traver 1959: ; – Elpers & Tomka 1994b: ; – Kluge 2004: ; – Zheng & Gong & Qiang & Zhou 2024:


Autapomorphy of Behningia/fg4.

(1) Larval hind leg [greatly modified – see Behningia/fg2 (1)] has femur greatly widened, with concavity into which tibia [greatly shortened – see Behningia/fg2 (1)] can be inputted. Tarsus is articulated to tibia movably, thus can bent at acute angle to tibia (in such a way that if leg would be non-specialized, tarsus would be directed outward). As a result, in folded condition femur, tibia and tarsus form a Z-figure, in which femur and tarsus are directed posteriorly. Patella-tibial suture of hind tibia is lost [but is retained on middle leg – see Behningia/fg2 (1)] (Kluge 2004: Fig.73:A–B).

Size. Fore wing length 15–24 mm. 

Distribution. Palaearctic and Oriental Region.


Nominal species in Behningia/fg3:  

baei McCafferty & Jacobus 2006 [Behningia]

 lestagei Motas & Bacesco 1937 [Behningia] ,,

 sushii Zheng & Zhou (in Zheng & Gong & Qiang & Zhou) 2024 [Behningia]

 tshernovae Edmunds & Traver 1959 [Behningia] ,,

 ulmeri Lestage 1930 [Behningia]

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