CLADOENDESIS OF EPHEMEROPTERA

ABC

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Anepeorus/fg1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata INCERTAE SEDIS)

Nomen hierarchicum: Anepeorus/fg1 [f:1962; g:1925] (incl. Spinadis)

In circumscription fits:

gen. Anepeorus McDunnough 1925: 190

gen. Spinadis Edmunds & Jensen 1974: 495

subfam. Spinadinae Edmunds & Jensen 1974: 495

Anepeorus/fg1: Kluge 2004: 209

Nominal taxon included: Spinadis/fg [fg:1974]


References. McDunnough 1925: ; Needham & Traver & Hsu 1935: ; Burks 1953: (non ); Edmunds & Jensen 1974 (Spinadis): ; Edmunds & Jensen & Berner 1976 (Anepeorus): (non ); ibid. (Spinadis): ; McCafferty & Provonsha 1984: ; 1985: ; 1986 (Spinadis): .


Characters of Anepeorus/fg1 of unclear phylogenetic status. 

(1) Mouth apparatus is adapted for carnivorism: Mandible is shortened perpendicular to axis of articulation, mola lacks grater. Superlinguae are vestigial. Maxilla lacks apical-ventral setae, its biting margin is shortened and both dentisetae are enlarged (McCafferty & Provonsha 1986: Fig.8,11,14). Non-unique apomorphy (see Index of characters [1.1.12]).

(2) Maxillary palp is 2-segmented (as in majority of other Pentamerotarsata, unlike Pseudiron).

(3) Larval paracercus is vestigial. Non-unique apomorphy (see Index of characters [1.3.64]); among Pentamerotarsata the same in Epeorus/fg1 and Acanthomola only.

Plesiomorphy of Anepeorus/fg1. Paraglossae are not widened (unlike Radulapalpata).

Size. Fore wing length 710 mm. 

Distribution. Nearctic.


Nominal species in Anepeorus/fg1:

 rusticus McDunnough 1925 [Anepeorus]

 simplex Walsh 1863 [Heptagenia]

 wallacei Edmunds & Jensen 1974 [Spinadis] syn.subj. simplex [Heptagenia]

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