CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Rhithrogena/fg4
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Branchitergaliae
Heptagennota
Pentamerotarsata
Radulapalpata
Rhithrogena/fg1Epeorus/fg1Rhithrogena/fg2CinygmulaRhithrogena/fg3
- Rhithrogena/fg4)
Nomen hierarchicum:
Rhithrogena/fg4
[f:1917; g:1881] (sine Epeiron,
Himalogena, Sibirigena)
In circumscription fits:
— Rhithrogena/fg4: Kluge 2004: 194
Non-typified taxa in Rhithrogena/fg4:
laevigata Tomka & Rash 1993
lobata Tomka & Rash 1993
Reference.
Kluge 2004: 
* 
*
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Characters of Rhithrogena/fg4 of unclear phylogenetic status. (1) In imago and subimago 1st segment of fore tarsus is short, several times shorter than 2nd. The same in Sibirigena, unlike Himalogena and Epeiron/g1; the same in some other taxa (see Index of characters [2.2.76]). (2) Each penis lobe is more or less tubular, with gonopore apically; sides near apex usually bear spines or denticles, including latero-dorsal [see Pentamerotarsata (5)] and medio-dorsal ones; titillators [ventral – see Rhithrogena/fg3 (5)] are always well-developed (Sowa 1970: Fig.1–21; Tomka & Hasler 1978: Fig.1–4). The same in Himalogena, unlike Sibirigena and Epeiron). |
Size. Fore wing length 5–17 mm.
Distribution. Holarctic.
Nominal species in Rhithrogena/fg4:
adrianae Belfiore 1983 [Rhithrogena]
allobrogica
Sowa & Degrange 1987 [Rhithrogena]
alpestris Eaton 1885 [Rhithrogena]
alpicola Navás 1935 [Rhithrogena]
— syn.subj.
alpestris [Rhithrogena]
austriaca Sowa & Weichselbaumer 1988 [Rhithrogena]
ayadi
Dakki & Thomas 1986 [Rhithrogena]
basalis [Baetis]
nomen nudum
— syn.subj. semicolorata [Baetis]
bajkovae
Sowa 1973 [Rhithrogena]
--/
beskidensis Alba-Tercedor
& Sowa 1987 [Rhithrogena]
binotata
Sinitshenkova 1982 [Rhithrogena]
— syn.subj. lepnevae [Rhithrogena]
bogoescui Sowa & Degrange 1987
[Rhithrogena]
braaschi Jacob 1974 [Rhithrogena]
brenneriana Klapálek
1905 [Rhithrogena]
— syn.subj. alpestris [Rhithrogena]
brodskyi Kustareva1976 [Rhithrogena]
brunneotincta
McDunnough 1933 [Rhithrogena]
bulgarica
Braasch & Soldán & Sowa 1985 [Rhithrogena]
buresi
Sowa 1973 [Rhithrogena]
carpatica Sowa 1971 [Rhithrogena picteti]
carpatoalpina Klonowska & Olechowska
& Sartori & Weichselbaumer 1987 [Rhithrogena]
castellana
Navás 1927 [Rhithrogena]
caucasica Braasch 1979 [Rhithrogena]
--/
cincta Navás 1921 [Rhithrogena]
circumtatrica Sowa
& Soldán 1986 [Rhithrogena]
colmarsensis Sowa 1984 [Rhithrogena]
— syn.subj. dorieri [Rhithrogena]
comitissa
Navás 1933 [Rhithrogena]
— syn.subj. gorrizi [Rhithrogena]
corcontica Sowa & Soldán 1986 [Rhithrogena]
dagestanica
Braasch 1979 [Rhithrogena]
daterrai Sowa 1984 [Rhithrogena]
decolorata Sinitshenkova 1973 [Rhithrogena]
degrangei Sowa
1969 [Rhithrogena]
delphinensis Sowa & Degrange 1987 [Rhithrogena]
diaphana Navás 1917 [Rhithrogena]
diensis Sowa
& Degrange 1987 [Rhithrogena]
dorieri
Sowa 1971 [Rhithrogena]
eatoni Esben-Petersen 1912 [Rhithrogena]
endenensis Metzler
& Tomka & Zurwerra 1985 [Rhithrogena]
— syn.subj. puthzi [Rhithrogena]
excisa
Sinitshenkova 1979 [Rhithrogena]
expectata Braasch 1979 [Rhithrogena]
--/
fasciata Traver 1933 [Rhithrogena]
ferruginea Navás 1905 [Rhithrogena]
fiorii Grandi
1953 [Rhithrogena]
fonticola Sowa & Degrange 1987 [Rhithrogena]
formosicola
Matsumura 1931 [Rhithrogena]
— syn.subj. parvus [Ecdyonurus]
fradleyi
Blair 1929 [Rhithrogena]
— syn.subj. germanica [Rhithrogena]
germanica Eaton 1885 [Rhithrogena]
giudicellorum
Thomas & Bouzidi 1986 [Rhithrogena]
goeldlini Sartori &
Sowa 1988 [Rhithrogena]
gorganica Klapálek 1907 [Rhithrogena]
gorrizi Navás 1913 [Rhithrogena]
gratianopolitana Sowa &
Degrange & Sartori 1986 [Rhithrogena]
grischuna Sartori &
Oswald 1988 [Rhithrogena]
grisoculata Bogoescu 1933 [Rhithrogena]
— syn.subj. semicolorata [Baetis]
haarupi Esben-Petersen 1909 [Rhithrogena]
— syn.subj. germanica [Rhithrogena]
henschii Klapálek 1906 [Rhithrogena]
hercegovina Tanasijevic 1985 [Rhithrogena]
hercynia Landa 1969 [Rhithrogena]
hybrida Eaton 1885 [Rhithrogena]
ingalik Randolph & McCafferty 2005 [Rhithrogena]
insularis
Esben-Petersen 1913 [Rhithrogena]
intermedia Zurwerra &
Metzler & Tomka 1987 [Rhithrogena]
iranica Braasch 1983 [Rhithrogena]
iridina
Kolenati 1859 [Baetis]
jacobi Braasch & Soldán 1988
[Rhithrogena]
jahorinensis Tanasievic 1985 [Rhithrogena]
japonica Ueno 1928 [Rhithrogena]
johannis Belfiore 1990 [Rhithrogena]
— syn.subj. siciliana [Rhithrogena]
joostiana
Sowa & Zimmermann 1976 [Rhithrogena]
--/
kimminsi
Thomas 1970
[Rhithrogena]
klausnitzeriana Braasch 1979 [Rhithrogena]
klugei Tiunova 2010 [Rhithrogena]
kownackii Sowa & Zimmermann 1976 [Rhithrogena iridina]
laciniosa
Sinitshenkova 1979 [Rhithrogena]
landai Sowa & Soldán
1984 [Rhithrogena]
lepnevae Brodsky 1930 [Rhithrogena]
--/
lisettae Baeuernfeind
2003 [Rhithrogena]
loyolaea Navás 1922 [Rhithrogena]
lucida
Braasch 1979 [Rhithrogena]
marcosi Alba-Tercedor & Sowa 1987
[Rhithrogena]
mariae
Vitte 1991 [Rhithrogena]
mariaedominicae
Sowa & Degrange 1987 [Rhithrogena]
marinkovici Tanasijevic
1985 [Rhithrogena]
monserrati Alba-Tercedor & Sowa 1986 [Rhithrogena]
neretvana Tanasijevic 1985 [Rhithrogena]
nivata Eaton 1871
[Heptagenia]
nuragica Belfiore 1987 [Rhithrogena]
orientalis You 1990 [Rhithrogena]
oscensis
Navás 1927 [Rhithrogena]
ourika
Thomas & Mohati 1985 [Rhithrogena]
parvus Ulmer 1912 [Ecdyonurus]
picteti
Sowa 1971 [Rhithrogena]
piechockii Braasch 1977 [Rhithrogena]
podhalensis Sowa & Soldán 1986 [Rhithrogena]
pontica
Sowa & Soldán & Kazanci 1986 [Rhithrogena]
potamalis
Braasch 1979 [Rhithrogena]
puthzi
Sowa 1984 [Rhithrogena]
puytoraci Sowa & Degrange 1987 [Rhithrogena]
quadrinotata
Sinitshenkova 1982 [Rhithrogena]
— syn.subj. bajkovae
[Rhithrogena]
reatina
Sowa & Belfiore 1984 [Rhithrogena]
robusta Dodds 1923 [Rhithrogena]
rolandi Weichselbaumer
1995 [Rhithrogena]
rubicunda Traver 1937 [Rhithrogena]
ryszardi
Thomas & Vitte & Soldán 1987 [Rhithrogena]
sangangensis You 1990 [Rhithrogena]
sartorii Zrelli & Boumaiza (in Zrelli & Sartori &
Bejaoui & Boumaiza) 2011 [Rhithrogena]
savoiensis Alba-Tercedor & Sowa 1987 [Rhithrogena]
/
semicolorata Curtis 1834 [Baetis]
— typus nominis Rhithrogena
/
semitincta
Pictet 1843 [Baetis]
— syn.subj. semicolorata [Baetis]
sibillina Metzler
& Tomka & Zurwerra 1985 [Rhithrogena]
siciliana Braasch 1989 [Rhithrogena]
soteria Navás
1917 [Rhithrogena]
sowai
Puthz 1972 [Rhithrogena]
strenua
Thomas (in Thomas & Gazagnes) 1982 [Rhithrogena]
sublineata Kazanci & Braasch
1988 [Rhithrogena]
tatrica Zelinka 1953 [Rhithrogena]
taurisca
Bauernfeind 1992 [Rhithrogena]
teberdensis Zimmermann 1977 [Rhithrogena]
--/
theischingeri
Braasch 1981 [Rhithrogena]
thomasi Alba-Tercedor & Sowa 1986
[Rhithrogena]
thracica Sowa & Soldán & Braasch
1988 [Rhithrogena]
trispina Zhou &
Zheng 2000 [Rhithrogena]
umbrosa
Braasch 1979 [Rhithrogena]
unicolor Tshernova 1952 [Rhithrogena]
— syn.subj. lepnevae [Rhithrogena]
ussingi Esben-Petersen 1910 [Rhithrogena]
— syn.subj. germanica [Rhithrogena]
vaillanti
Sowa & Degrange 1987 [Rhithrogena]
vulpecula Klapálek 1905 [Rhithrogena]
— syn.subj. semicolorata [Baetis]
wolosatkae Klonowska 1987
[Rhithrogena]
zelinkai Sowa & Soldán 1984 [Rhithrogena]
zernyi
Bauernfeind 1991 [Rhithrogena]
zhiltzovae Sinitshenkova
1979 [Rhithrogena]
See also:
Rhithrogena/fg3
INCERTAE SEDIS
Radulapalpata
INCERTAE SEDIS
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