CLADOENDESIS OF EPHEMEROPTERA |
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Paegniodes
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Branchitergaliae
Heptagennota
Pentamerotarsata
Radulapalpata
Rhithrogena/fg1
-
Paegniodes)
Nomen hierarchicum: Paegniodes/g(1) [g:1881]
In circumscription fits:
— subgen. Paegniodes: Kluge 1988: 305
— gen. Paegniodes Eaton 1881: 23
— Paegniodes/g(1): Kluge 2004: 190
References. Eaton 1883–1888: ; – Ulmer 1925: ; – 1936: ; – Hsu 1932: ; – 1936: ; – Ueno 1969: ; – Tshernova 1976: ; – Kluge 1988: ; – 1993: ; – 2004: .
Plesiomorphies of Paegniodes. Unlike other Rhithrogena/fg1 (i.e. Rhithrogena/fg2 and Epeorus/fg1), subimaginal lateroparapsidal stripe of pigmented area of mesonotum retains shape initial for Heptagennota [see Heptagennota (2)], i.e. is widened toward apex and its posterior-median angle touches medioparapsidal suture [in spite of the fact that lateroparapsidal suture is curved and does not touch medioparapsidal suture – see Rhithrogena/fg1 (3)] (Kluge 2004: Fig.64:A). Unlike Rhithrogena/fg2: maxilla is long [see Radulapalpata (3)], ventral row of setae [see Branchitergaliae (1)] is straight, parallel to median margin (Kluge 1988: Fig.38) (the same in Cinygma and Epeorus/fg1 – see Index of characters [1.1.30]); penis without additional pair of ventral titillators, with a pair of well-developed median titillators [see Pentamerotarsata (5)]; apices of these median titillators project at ventral side of penis being directed laterally, thus have some resemblance with a single pair of ventral titillators in Rhithrogena/fg3; but unlike ventral titillators, they arise from median walls of penis lobes. Inlike Epeorus/fg1: larval paracercus is as long as cerci, primary swimming setae [vestigial – see Pentamerotarsata (6)] are present; imaginal and subimaginal mesonotal suture is normally developed [see Plesiomorphies of Branchitergaliae] (Kluge 2004: Fig.64:A). Each superlingua is roundish, not narrowing toward apex (Tshernova 1976: Fig.43) (unlike Rhithrogena/fg2 and Epeorus/fg3), but lateral margins of superlinguae are not divergent (unlike Bleptus, Ironodes and other Pentamerotarsata [for comment – see Ironodes (3) below]. Glossae are pyramidal [see Radulapalpata (1)] (Tshernova 1976: Fig.45). Larval and adult patella-tibial suture is developed on middle and hind legs (unlike most of larval Epeorus/fg1), absent on fore legs (unlike larval Rhithrogena/fg3). |
Size. Fore wing length 10–20 mm.
Distribution. Oriental Region.
Nominal species in Paegniodes/g(1):
cupulata Eaton 1871 [Heptagenia] — typus nominis Paegniodes -,/,,
dao Nguyen & Bae 2004 [Paegniodes] — syn.subj. cupulata [Heptagenia]
fukiensis Hsu 1932 [Paegniodes] — syn.subj. cupulata [Heptagenia]
oligopilus Shi & Tong 2023 [Paegniodes]
sapanensis Boonsoong & Sartori & Auychinda (in Boonsoong & Auychinda & Sartori & Khanyom) 2021 [Paegniodes]
See also:
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