CLADOENDESIS OF EPHEMEROPTERA |
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Ironodes
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Branchitergaliae
Heptagennota
Pentamerotarsata
Radulapalpata
Rhithrogena/fg1
Epeorus/fg1 BleptusEpeorus/fg2
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Ironodes)
Nomen hierarchicum: Ironodes/g(1) [g:1935]
In circumscription fits:
— subgen. Ironodes: Kluge 1988: 308
— gen. Ironodes Traver 1935: 32
— Ironodes/g(1): Kluge 2004: 202
References. Traver 1935: * *; – Needham & Traver & Hsu 1935: * *; – Day 1952: '; – Edmunds & Jensen & Berner 1976: * *; – Tshernova 1976: *; – Kluge 1988: *; – 2004: * *.
Autapomorphies of Ironodes. (1) Tergalius VII lacks fibrillose lobe [see Branchitergaliae (3)]. Non-unique apomorphy; among Radulapalpata the same in some Heptagenia/f5=g4 and some Cinygmula. (2) Titillators [see Pentamerotarsata (5)] are lost. Non-unique apomorphy; among Epeorus/fg1 the same in Bleptus and some Epeorus/fg3 (see Index of characters [2.3.15]). Character of Ironodes of unclear phylogenetic status. (3) Superlinguae diverge toward apex (Kluge 1988: Fig.29). The same in Bleptus (ibid.: Fig.96). Probably, this is a symplesiomorphy of Bleptus and Ironodes within Epeorus/fg1, but in this case similarity of Rhithrogena/fg2 and Epeorus/fg3 (whose superlinguae are widest in middle part and convergent toward apex) can be explained only as a convergention. Possibly, vice verse, convergention is the similarity of Bleptus and Ironodes, while similarity of Rhithrogena/fg2 and Epeorus/fg3 is a symplesiomorphy within Rhithrogena/fg1. Plesiomorphies of Ironodes. Mandible is longer than in Epeorus/fg2, with incisor and kinetodontium slender. Maxilla is long [see Radulapalpata (3)], apical margin bears long hairs and vestiges of pectinate setae [see Epeorus/fg1 (5)], canines are not thickened (Kluge 1988: Fig.34–35) (symplesiomorphy with Bleptus, unlike Epeorus/fg3). Larval cerci have no dorsal hairs (unlike Epeorus/fg3). |
Size. Fore wing length 8–14 mm.
Distribution. Western Nearctic.
Nominal species in Ironodes/g(1):
californicus Banks 1910 [Heptagenia (Epeorus)]
flavipennis Traver 1935 [Ironodes]
geminatum Eaton 1885 [Cinygma]
lepidus Traver 1935 [Ironodes]
nitidus Eaton 1885 [Iron] — typus nominis Ironodes
See also:
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