CLADOENDESIS OF EPHEMEROPTERA |
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Drunella/g1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Furcatergaliae
Ephemerella/fg1Tricorythus/fg1Ephemerella/fg2
Timpanoga/fg1Ephemerella/fg3
Torleya/g1Ephemerella/fg4
-
Drunella)
Nomen hierarchicum: Drunella/g1
[g:1905]
In circumscription fits:
— subgen. Drunella: McDunnough 1931b: 210
— gen. Drunella: Allen 1980: 79
— Drunella/g1: Kluge 2004: 304
Nominal taxa included:
References.
Edmunds 1959: *
*;
– Allen & Edmunds 1962a:
*
*; – Edmunds &
Jensen & Berner 1976:
*
*; – Kluge 2004:
*
*
*
*
Autapomorphies of Drunella. (1) Larva is carnivorous, fore legs are specialized as grasping: fore femur is usually thickened, usually widest in its proximal half; transverse row of setae [see Ephemerella/fg1 (12)] is always absent; inner (directed anteriorly) margin of fore femur is usually serrate, that allows to keep a prey, clutching it between the femur and tibia; fore tibia on its apex on inner side has a projection directed distally, along inner side of tarsus (Kluge 1997d: Fig.18.4–18.8). In selected representatives of Drunella/fg1 some of these characters are not expressed or weakly expressed: fore femur can be nearly parallel-sided, and its denticles can be the same as on middle and hind femora or absent. Among the characters named here, the most constant is the projection on tibia. Besides Drunella/g1, similar specialization of fore femur (but not tibia) independently appeared in Tricorythus/fg1-Vietnamella. (2) Larval clypeus is crossed transversely by a distinct stripe of hairs projected forward (Allen 1973: Fig.8). Unique apomorphy: some other Ephemerella/fg1 have similar transverse row of setae on frons, but not on clypeus. Plesiomorphies of Drunella. Maxilla is biting [see Ephemerella/fg1 (16)], vestige of maxillary palp is always present, 3-segmented [see Ephemerella/fg1 (1)]. Larval thorax without projections characteristic for Cincticostella/g1 or Notacanthella. Larval paracercus has the same length as cerci (unlike Caudatella). |
Size. Fore wing length 6–19 mm.
Distribution. Amphipacific: Nearctic and Eastern Palaearctic; the most western representatives inhabit Tien-Shan-Himalayan Region (all former reports of Drunella from more western areas appeared to be wrong).
Nominal species in Drunella/g1:
aculea Allen 1971 [Ephemerella (Drunella)]
-
-
/
allegheniensis Traver 1934 [Ephemerella]
autumnalis McDunnough 1934 [Ephemerella]
— syn.subj. spinifera
[Ephemerella]
basalis Imanishi 1937 [Ephemerella]
bicornis Gose 1980 [Ephemerella
(Drunella)]
— syn.subj. lepnevae [Ephemerella]
bifurcata Allen 1971 [Ephemerella (Drunella)]
— syn.subj. sachalinensis [Ephemerella]
bispina Needham 1905 [Ephemerella]
— syn.subj. walkeri [Ephemerella]
borakensis Allen 1971 [Ephemerella
(Drunella)]
— syn.subj. submontana [Ephemerella]
carsona
Day 1952 [Ephemerella glacialis]
cherokee Traver 1937 [Ephemerella]
— syn.subj. tuberculata [Ephemerella]
coloradensis Dodds
1923 [Ephemerella]
— typus nominis Myllonella
,
-
conestee
Traver 1932 [Ephemerella]
— syn.subj. tuberculata [Ephemerella]
cornuta
Morgan 1911 [Ephemerella]
— syn.subj.
lata [Ephemerella]
cornutella
McDunnough 1931 [Ephemerella]
cryptomeria Imanishi 1937 [Ephemerella]
depressa Ide 1930 [Ephemerella]
— syn.subj.
lata [Ephemerella]
doddsi Needham 1927 [Ephemerella (Eatonella)]
— typus nominis Eatonella
,
ezoensis
Gose 1980 [Ephemerella (Drunella) trispina] nom.praeocc.
— syn.subj. triacantha [Ephemerella]
flavilinea
McDunnough 1926 [Ephemerella]
flavitincta
McDunnough 1934 [Ephemerella]
— syn.subj. grandis [Ephemerella]
fuscata
Walker 1853 [Baetis] nom. praeocc.
— syn.obj. walkeri [Ephemerella]
fuso
Jacobus & McCafferty 2008 [Drunella]
— syn.obj. fusongensis [Drunella]
fusongensis Su & Gui 1995 [Drunella]
— syn.subj. lepnevae [Ephemerella]
glacialis Traver
1934 [Ephemerella]
— syn.subj. grandis [Ephemerella]
grandis Eaton 1884 [Ephemerella]
— typus nominis Drunella
,
inflata
McDunnough 1926 [Ephemerella]
— syn.subj.
lata [Ephemerella]
ingens
McDunnough 1934 [Ephemerella]
— syn.subj. grandis [Ephemerella]
kabulensis Allen 1973 [Ephemerella (Drunella)]
— syn.subj. submontana [Ephemerella]
kohnoi
Allen 1971 [Ephemerella (Drunella)]
lapidula McDunnough
1935 [Ephemerella]
— syn.subj. flavilinea
[Ephemerella]
lata Morgan 1911 [Ephemerella]
latipes Tshernova 1952 [Ephemerella]
-
-
/
lepnevae Tshernova 1949 [Ephemerella]
-
-
/
longicornis Traver 1932 [Ephemerella]
— syn.subj.
lata [Ephemerella]
longipes
Tshernova 1952 [Ephemerella]
— syn.subj. lepnevae [Ephemerella]
nasiri Ali 1971 [Ephemerella]
— syn.subj. submontana [Ephemerella]
pelosa
Mayo 1951 [Ephemerella]
proserpina Traver 1934 [Ephemerella]
— syn.subj. grandis [Ephemerella]
sachalinensis Matsumura 1931 [Ephemerella]
sierra
Mayo 1952 [Ephemerella]
— syn.subj. spinifera
[Ephemerella]
solida
Bajkova 1980 [Ephemerella]
-
-
/
spinifera Needham 1927
[Ephemerella (Drunella)]
submontana Brodsky 1930 [Ephemerella]
-
-
/
svenhedini
Ulmer 1936 [Ephemerella]
— syn.subj. submontana [Ephemerella]
tenax Tshernova 1952 [Ephemerella]
— syn.subj. triacantha [Ephemerella]
traverae Allen & Edmunds 1963 [Ephemerella
(Drunella)]
— syn.subj. submontana [Ephemerella]
triacantha Tshernova
1949 [Ephemerella]
-
-
/
trispina Ueno
1928 [Ephemerella]
—
typus nominis Tribrochella
tuberculata Morgan 1911 [Ephemerella]
—
typus nominis Unirachella
walkeri Eaton
1884 [Ephemerella]
wayah Traver 1932 [Ephemerella]
— syn.subj. walkeri [Ephemerella]
wilsoni Mayo 1952 [Ephemerella]
— syn.subj. coloradensis [Ephemerella]
yosemite
Traver 1934 [Ephemerella]
— syn.subj. grandis [Ephemerella]
yoshinoensis Gose 1963 [Ephemerella]
See also:
Ephemerella/fg3
INCERTAE SEDIS
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