CLADOENDESIS OF EPHEMEROPTERA

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Caucasiron

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata  Rhithrogena/fg1  
Epeorus/fg1
BleptusEpeorus/fg2IronodesEpeorus/fg3 Iron/g1 Ironopsis/g1 - Caucasiron)

Nomen hierarchicum: Caucasiron/g(1) [g:1997]

In circumscription fits:

— subgen. Caucasiron Kluge 1997a: 233

— Caucasiron/g(1): Kluge 2004: 207


References. Tshernova 1938 (Cinygma caucasica): '; – Sinitshenkova 1976 (Iron): ' '; – Braasch 1978 (Iron): '; – 1979 (Iron): ' '; – Braasch & Soldan 1979 (Iron): '; – Kluge 1997a: * *; – 1997d: *; – 2004: * * *.


Autapomorphy of Caucasiron.

(1) Each tergalius II–VII on its costal rib [which is situated on costal margin, bears spines and setae – see Epeorus/fg3 (2)] has a sharp projection covered with spines and directed anteriorly-dorsally (Kluge 2004: Fig.65:G); this projection can be well-developed on all tergalii II–VII, or at least on anteriormost of them. Unique apomorphy.

Plesiomorphy of Caucasiron. Penis has a pair of well-developed median titillators [see Pentamerotarsata (5)] (unlike Ironopsis/g2, Epeorus/fg4 and some others).

Variable characters of Caucasiron. Imaginal and subimaginal claws are usually ephemeropteroid, but in guttatus [Iron] – pointed (see Variable characters of Heptagennota and Index of characters [2.2.85]).

Size. Fore wing length 10–20 mm.

Distribution. Greece, Caucasus and Tienshan-Himalayan Rhithrofaunisctic Region.


Nominal species in Caucasiron/g(1):

 abditus Hrivniak & Sroka (in Hrivniak & Sroka & Türkmen & Martynov  & Bojkova) 2024 [Epeorus (Caucasiron)]

  alborzicus Hrivniak & Sroka (in Hrivniak & Sroka & Bojková & Godunko & Namin  & Bagheri & Nejat & Abdoli & Staniczek) 2020 [Epeorus (Caucasiron)]

 alpestris Braasch 1979 [Iron]

bicolliculatus Hrivniak (in Hrivniak & Sroka & Godunko & Žurovcovà) 2017 [Epeorus (Caucasiron)]

 caucasica Tshernova 1938 [Cinygma] — typus nominis Caucasiron --/

extraordinarius Chen & Wang & Zhou 2010 [Epeorus (Caucasiron)]

 fuscus Sinitshenkova 1976 [Iron]

 guttatus Braasch & Soldán 1979 [Iron] --/

 insularis Braasch 1983 [Iron znojkoi]

 iranicus Braasch & Soldan 1979 [Iron caucasicus]  

 longimaculatus Braasch 1980 [Iron]

 magnus Braasch 1978 [Iron] --/

 nigripilosus Sinitshenkova 1976 [Iron]

psi Eaton 1885 [Epeorus] ,

 shargi Hrivniak & Sroka (in Hrivniak & Sroka & Bojková & Godunko & Namin  & Bagheri & Nejat & Abdoli & Staniczek) 2020 [Epeorus (Caucasiron)]

 sinitshenkovae Braasch & Zimmermann 1979 [Iron]

 soldani Braasch 1979 [Iron]

suspicatus Braasch 2006 [Iron]

 tripertitus Hrivniak & Sroka (in Hrivniak & Sroka & Godunko & Manko & Bojkova) 2022 [Epeorus (Caucasiron)]

 turcicus Hrivniak & Sroka & Türkmen & Godunko & Kazanci 2019 [Epeorus (Caucasiron)]

 zagrosicus Hrivniak & Sroka (in Hrivniak & Sroka & Bojková & Godunko & Namin  & Bagheri & Nejat & Abdoli & Staniczek) 2020 [Epeorus (Caucasiron)]

 znojkoi Tshernova 1938 [Iron] --/


Examined also:

 spp.


See also:

Epeorus/fg3 INCERTAE SEDIS

Radulapalpata INCERTAE SEDIS

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