CLADOENDESIS OF EPHEMEROPTERA |
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Anagenesia/g1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Furcatergaliae
Fimbriatotergaliae
Fossoriae
Cryptoprosternata
Palingenia/f2=g1PentageniaPalingenia/f3=g2
- Anagenesia/g1)
Nomen hierarchicum: Anagenesia/g1 [g:1883] (incl. Mortogenesia, Chankagenesia, Plethogenesia, Cheirogenesia)
In circumscription fits:
— subgen. Anagenesia Eaton 1883–1888: 25
— gen. Anagenesia: Lestage 1917: 256
— Anagenesia/g1: Kluge 2004: 254
References. Eaton 1883–1888: *; – Kluge 2004: * * * *
Autapomorphies of Anagenesia/g1. (1) Fore wing is modified in following manner: RSa2 [approximated to iRS – see Palingenia/f3=g2 (2)] does not form a triad (non-unique apomorphy – see Index of characters [2.2.37]); bifurcation of MA is transferred to proximal part of wing (non-unique apomorphy – see Index of characters [2.2.43]); additional intercalaries are absent or weekly developed, thus anterior-distal part of wing is armed only by paired veins RSa2+iRS, RSp+MA1 and iMA+MA2 [see Palingenia/f3=g2 (2)], wing margin between their apices can be incised (Kluge 2004: Fig.75:A; Demoulin 1965b: Fig.3–8). This form of wing has convergent similarity with Behningia/fg3. Some representatives of Anagenesia/g1 retain ability to fly, while some of them (Chankagenesia, Cheirogenesia, partly Anagenesia/g2) are unable to fly and can only skim the water surface (Buldovsky 1935a, 1935b; Tshernova 1952; Sartori & Elouard 1999). (2) In male and female imago, legs [non-functional – see Palingenia/f3=g2 (1)] are strongly modified. In male imago fore leg (elongate in majority of mayflies) retains subimaginal structure [see Palingenia/f3=g2 (3)]: it is short, with tarsus strongly shortened, thickened, usually arched (Demoulin 1965b: Fig.3–8). In both sexes on middle and hind legs all claws are blunt; in male posterior claw (i.e. the claw corresponding to pointed claw when claw structure is ephemeropteroid) can be more or less reduced (Table) [claws of fore leg are also blunt – see Palingenia/f2=g1 (7) and Palingenia/f3=g2 (4)]. Female legs are reduced in greater degree; not only fore tarsus [see Palingenia/f3=g2 (4)], but all tarsi are non-segmented, each with 1 blunt claw only. Male subimago [skimming the water surface – see (1)] can moult to imago on the water surface immediately after moult from larva to subimago, some specimens do not moult at all [females never moult – see Palingenia/f3=g2 (1)] (Buldovsky 1935a, 1935b). Some of these features independently appeared in some other short-living mayflies. (3) Vestige of tergalius I [initially bilamellate – see Cryptoprosternata (6)] is unilamellate. Non-unique apomorphy (see Index of characters [1.3.33]). |
Size. Fore wing length 11–30 mm.
Distribution. Asia, New Guinea, Madagascar.
The taxon Anagenesia/g1 is divided into: |
These taxa differ one from another by combinations of non-unique characters:
Taxa |
Geographical distribution |
Species number |
Characters |
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Imaginal |
Larval |
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1 |
2 |
3 |
4 |
5 |
6 |
7 |
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Palingenia/f4=g3 |
West Palaearctic |
6 |
I |
I I |
0 ) |
4–6 |
Δ |
Δ |
2 |
Anagenesia/g1: |
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Mortogenesia |
Iraq |
1 |
I |
I I |
00 |
4–6 |
Δ |
Δ |
1 |
Chankagenesia |
Eastern Asia |
1 |
I |
I I |
0 |
2 |
I |
I |
1 |
Anagenesia/g2 |
Asia |
14 |
Y |
I I |
0 |
2–4 |
I |
I |
1 |
Plethogenesia |
New Guinea |
4 |
Y |
I I |
00 |
2 |
Δ |
I |
1 |
Cheirogenesia |
Madagascar |
1 |
Y |
II |
00 |
2 |
I |
I |
1 |
1 – vein CuA of fore wing: "I" – simple, "Y" – with bifurcation
2 – veins MP1 and iMP of fore wing: "II" – brought together, "I I" – not brought together
3 – claws of middle and hind legs of male imago: "0 )" – two ephemeropteroid, "00" – two blunt, "0" – single blunt
4 – number of segments of gonostylus
5 – tusk denticles: " I " – denticle with articulated apical spine-like seta; "Δ" – integral denticle
6 – outer margin of fore tibia of larva: " I " – with spine-like setae; "Δ" – with denticles
7 – number of lamellae of first tergalius