CLADOENDESIS OF EPHEMEROPTERA

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Plesiomorphon Rhithrogeniella/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata
Heptagenia/f5=g4 
Ecdyonurus/fg1 - pm.Rhithrogeniella/g1)

Nomen hierarchicum: Rhithrogeniella/g1 [g:1964].

In circumscription fits:

— gen. Nixe Flowers 1980: 299

— gen. Afghanurus: Braasch 1981: 128

— plesiomorphon Afghanurus/g2: Kluge 2004: 180

— subgen. Rhithrogeniella: Kluge & Srinivasan  & Sivaruban & Barathy & Isack 2023: 501-523.: 503

Nominal taxa included:

Afghanurus/g [g:1964]  

Paracinygmula/g [g:1975] 

Nixe/g [g:1980] 

Akkarion/g [g:1980]


Reference. Flowers 1980: * *; – Kluge 2004: * * *; – Kluge & Srinivasan  & Sivaruban & Barathy & Isack 2023: 501-523..


Characters of Rhithrogeniella/g1 of uncleart phylogenetic status.

(1) Ventral sclerites of penis distad-laterad of titillators [see Pentamerotarsata (5)], bear one or several pairs of denticles ("discal spines") directed distally-laterally (Kluge 1980: Fig.64,71,83,90,103). Only in a few species these "discal spines" are secondarily lost (particularly in ornata Rhithrogeniella] – Kluge et al. 2023: Figs 69–73). The same in Compsoneuriella/g1.

Plesiomorphies of Rhithrogeniella/g1.

(2) Maxilla with distal dentiseta simple, non-branched (Kluge et al. 2023: Fig. 33) (the same in Ecdyonurus/fg, Leucrocuta and Ecdyogymnurus; in contrast to Atopopus/fg1).

(2) Larval pronotum without lateral projections stretched laterad of mesonotum (Kluge et al. 2023: Figs 1–3) (in contrast to Ecdyonurus/fg2).

(4) Larval caudalii with swimming setae: regular or irregular row of primary swimming setae is present at least on each lateral side of paracercus and on median (i.e. inner) side of each cercus (Kluge et al. 2023:Fig. 47) (the same in Ecdyonurus/fg2, in contrast to other taxa within Ecdyonurus/fg1). Similar secondary setae on lateral (i.e. outer) side of each cercus are either absent, or present: they are present in all examined individuals of ornatus [Rhithrogeniella] (Kluge et al. 2023: Figs 48–50) and in some individuals of joernensis [Ecdyonurus] (Kluge et al. 2023: Fig. 52), but absent in other examined species.

Size. Fore wing length 5–9 mm.

Distribution. Holarctic (dominate in Eastern Palaearctic and Nearctic).


Nominal species in Rhithrogeniella/g1:

bajkovae Kluge (in Tshernova & Kluge &  Sinitshenkova & Belov) 1986 [Ecdyonurus] — syn.obj. zhiltzovae [Paracinygmula]

criddlei McDunnough 1927 [Heptagenia]

dentata Braasch 1979 [Heptagenia] — syn.subj. joernensis [Ecdyurus]

flavomaculatus Aro 1928 [Ecdyonurus] — syn.subj. joernensis [Ecdyurus]

imanica Bajkova 1972 [Rhithrogena] — syn.subj.  simplicioides [Heptagenia]

inconspicua McDunnough 1924 [Heptagenia]

joernensis Bengtsson 1909 [Ecdyurus] --/

littorosus Kang & Yang 1994 [Nixe (Nixe)]

lucidipennis Clemens 1913 [Ecdyurus] — typus nominis Nixe

mitificus Kang & Yang 1994 [Nixe (Nixe)]

mongolica Bajkova & Varykhanova 1978 [Heptagenia] — syn.subj. joernensis [Ecdyurus]

obscurus Kang & Yang 1994 [Nixe (Nixe)]

ornata Ulmer 1939 [Rhithrogeniella] — typus nominis Rhithrogeniella --/

otiosa McDunnough 1935 [Heptagenia] — syn.subj. criddlei [Heptagenia]

perfida McDunnough 1926 [Heptagenia]

inconspicua McDunnough 1924 [Heptagenia]

rodocki Traver (in Needham & Traver & Hsu) 1935 [Heptagenia] — syn.subj. simplicioides [Heptagenia]

rubrofasciatus Brodsky 1930 [Ecdyonurus] --/

rusticalis McDunnough 1931 [Heptagenia]

simplicioides McDunnough 1924 [Heptagenia] --/

stubbei Braasch 1979 [Ecdyonurus] — syn.subj. joernensis [Ecdyurus]

subspinosa Braasch & Soldán 1988 [Nixe] — syn.subj. zhiltzovae [Paracinygmula]

tianshanica Kustareva 1984 [Stenonema] — syn.subj. rubrofasciatus [Ecdyonurus]

 tonkinensis Soldán & Braasch 1986 [Rhithrogeniella] — syn.subj. ornata [Rhithrogeniella]

vicinus Demoulin 1964 [Afghanurus] --/

werestschagini Tshernova 1952 [Heptagenia] — syn.subj. simplicioides [Heptagenia]

zhiltzovae Bajkova 1975 [Paracinygmula] --/


Examined also:

sp.n. (south India) -/


See also:

Ecdyonurus/fg1 INCERTAE SEDIS

Radulapalpata INCERTAE SEDIS

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