EPHEMEROPTERA OF THE WORLD

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sources:

subjects:

valid names:

invalid synonyms:

Müller 1776

Dania

Ephemera sulphurea sp.n.

Pictet 1843-1845

Baetis sulphurea comb.n.

?= Ephemera helvola syn.n.

?= Baetis straminea syn.n.

Walker 1853

Baetis sulphurea

?= Ephemera helvola 

?= Baetis straminea

Brauer 1857:74

Baetis sulphurea

Ausser. 1869

Baetis sulphurea

Eaton 1871*

syn. Heptagenia elegans syn.n.

Joly 1876

anatom.

Baetis sulphurea

Eaton 1883-1888

variation

Heptagenia sulphurea comb.n.

= Ephemera helvola

= Baetis straminea

= Ephemera leucophthalma syn.n.

= Ephemera ferruginea syn.n.

= Ephemera citrina syn.n.

= Baetis elegans syn.n.

= Baetis costalis syn.n.

?= Baetis marginalis syn.n.

= Baetis cyanops syn.n.

= Baetis lutea:
Hagen 1863
(non Linnaeus 1767)

= Ephemera bioculata:
Römer 1789
Panzer 1804 (var.)
Panzer 1805
(non Linnaeus 1758)

Eaton 1887'

Portugal

Heptagenia sulphurea

Rostock (1887)1888

Heptagenia sulphurea

Mocsary 1899

Heptagenia sulphurea

Eaton 1901

Norvegia

Heptagenia sulphurea

Esben-Petersen 1910

Heptagenia sulphurea

Neeracher 1910

Heptagenia sulphurea

Leue 1911

anat.

Heptagenia sulphurea

! Lestage 1919

Bengtsson 1913

Heptagenia sulphurea

Pongracz 1914

Heptagenia sulphurea

Bngtsson 1917

Heptagenia sulphurea

Navas 1917:81-91

España

Heptagenia sulphurea

Lestage 1917

Heptagenia sulphurea

Lestage 1919

Heptagenia sulphurea

sensu Leue 1911
= Ecdyurus fusco-griseus

Ulmer 1920:97-144*

-

Heptagenia sulphurea

Navas 1925:157-161'

pen.Iberica

Heptagenia sulphurea

Tshernova 1928

r.Oka

Heptagenia sulphurea

Percival & Whitehead 1929

Heptagenia sulphurea

Ulmer 1929

Heptagenia sulphurea

Schoenemund 1930

Heptagenia sulphurea

Schoenemund 1930

España

Heptagenia sulphurea

Neizvestnova-Zhadina 1930

r.Oka

Heptagenia sulphurea

Kolesov 1930*

Heptagenia sulphurea

Meyer E. 1931

blood system

Heptagenia sulphurea

Navas 1932

España

Heptagenia sulphurea

Eidel 1933*

Deutschland

Heptagenia sulphurea

Tiensuu 1935

Heptagenia sulphurea

-"-

Heptagenia sulphurea f. denervosa

Mackiewicz-Gutowska 1935*

Poland

Heptagenia sulphurea

Kimmins 1936:147-151

Britain

Heptagenia sulphurea

Mikulski 1936

Heptagenia sulphurea

Tshernova 1936

Moscow.prov.

Heptagenia sulphurea

Tshernova 1937

r.Dnepr

Heptagenia sulphurea

Tiensuu 1937

anomal.ven.

Heptagenia sulphurea

Eidel 1937*

Deutschland

Heptagenia sulphurea

Borang 1937

Estonia

Heptagenia sulphurea

Kimmins 1938:202-209

Britain

Heptagenia sulphurea

Brekke 1938*

Norvegia

Heptagenia sulphurea

Mosely 1938

oviposition

Heptagenia sulphurea

Beling 1939

r.Dnepr

Heptagenia sulphurea

Tiensuu 1939

Heptagenia sulphurea

Macan & Macan 1940

r.Avon

Heptagenia sulphurea

Satori & Nagy 1940

Heptagenia sulphurea

Zhadin 1940

Russia

Heptagenia sulphurea

Tshernova 1941

n.Russia

Heptagenia sulphurea

Kimmins 1942

Heptagenia sulphurea

Tshernova 1944*

Heptagenia sulphurea

Tshernova 1947

n.Russia

Heptagenia sulphurea

Real 1947

Heptagenia sulphurea f. cauzonica

Zelinka 1953:181-200

Heptagenia sulphurea

Tshernova 1953*

n.Russia

Heptagenia sulphurea

Pleskot 1953

Wienerwald

Heptagenia sulphurea

Haberman 1953*

Estonia

Heptagenia sulphurea

Macan 1955:59-70

Heptagenia sulphurea

Tabacaru 1956*

Romania

Heptagenia sulphurea

Degrange 1956*

micropyle

Heptagenia sulphurea

Jensen C.F. 1956

Denmark

Heptagenia sulphurea

Harris 1956

oviposition

Heptagenia sulphurea

Macan 1958*

Heptagenia sulphurea

Bogoescu 1958

Heptagenia sulphurea

Kazlauskas 1959

Lithuania

Heptagenia sulphurea

Ujhelyi 1959

Heptagenia sulphurea

Sowa 1959

Heptagenia sulphurea

Müller-Liebenau 1960

Eifel

Heptagenia sulphurea

Keffermuller 1960

Heptagenia sulphurea

Grandi 1960

Heptagenia sulphurea

Degrange 1960*

reprod., , 
raptor-ovi, larvula
parthen.

Heptagenia sulphurea

Landa 1962*

Heptagenia sulphurea

Macan 1961:1-64

Britain

Heptagenia sulphurea

Bogoscu & Tabacaru 1962*

Heptagenia sulphurea

Kazlauskas & Sanvaitite 1962

Latvia

Heptagenia sulphurea

Tshernova 1964*

Heptagenia sulphurea

Gerd & Sokolova 1965

Karelia

Heptagenia sulphurea

Ujhelyi 1966

Heptagenia sulphurea

Illies 1967

Heptagenia sulphurea

Ulfstrand 1968

Heptagenia sulphurea

Madsen 1968

Heptagenia sulphurea

Machel 1969

Poland

Heptagenia sulphurea

Landa 1969

Heptagenia sulphurea

Saftoiu 1969

neurosecret.

Heptagenia sulphurea

Langford 1971

Heptagenia sulphurea

Kachalova & Skrube 1971*

Latvia

Heptagenia sulphurea

Baba & Ferencz 1971

Heptagenia sulphurea

Jazdzewska 1971*

Heptagenia sulphurea

Bajkova 1972

Krasnojarsk

Heptagenia sulphurea

Kasymov 1972

Caucasus

Heptagenia sulphurea

Malzacher 1973*

Heptagenia sulphurea

Dahlby 1973

Norvegia

Heptagenia sulphurea

Müller-Liebenau 1973

DeutschDR

Heptagenia sulphurea

Jacob 1974

Heptagenia sulphurea

= Heptagenia marginalis 

Sowa 1975

Heptagenia sulphurea

Sowa 1975

Heptagenia sulphurea

Malzacher 1976*

Heptagenia sulphurea

Braasch & Jacob 1976*

Heptagenia sulphurea

Kazlauskas 1977

Heptagenia sulphurea

Illies 1978

Heptagenia sulphurea

Braasch 1979*

Mongolia

Heptagenia sulphurea

Fischer 1979

oculi

Heptagenia sulphurea

Soldan 1979:223-230*

spermatozoid

Heptagenia sulphurea

Puthz 1980

Albania

Heptagenia sulphurea

Saaristo & Savolainen 1980

Heptagenia sulphurea

≠ Heptagenia dalecarlica

Elliott & Humpesch 1980*

oviposition

Heptagenia sulphurea

Alba-Tercedor 1981*

España

Heptagenia sulphurea

Savolainen & Saaristo 1981*

Finland

Heptagenia sulphurea

Malzacher 1981:41-72*

s-w.Deutschland

Heptagenia sulphurea

Malzacher 1981:145-147*

rote liste

Heptagenia sulphurea

Sander 1981

Schwarzwald

Heptagenia sulphurea

Spuris 1982*

Latvia

Heptagenia sulphurea

Elliott & Humpesch 1983

Heptagenia sulphurea

Braasch 1983

Mongolia

Heptagenia sulphurea

Mol 1984:IV.Conf.Eph.

Netherlands

Heptagenia sulphurea

Jazdzewska 1984:IV.Conf.Eph.

Heptagenia sulphurea

Zelinka 1984:IV.Conf.Eph.

Heptagenia sulphurea

Gonzalez del Tanago 1984

España

Heptagenia sulphurea

Savolainen & Saaristo 1984

Finland

Heptagenia sulphurea

Zurwerra & Tomka 1984

Switzerland

Heptagenia sulphurea

Novikova 1984

n.Urals

Heptagenia sulphurea 

Kiseleva & Ezernitsky 1985

Crimea

Heptagenia sulphurea

! Prokopov  & Godunko 2007

Lallab-Wakim 1985*

(in detail)

growth

Heptagenia sulphurea

Sartori & Dethier 1985

Switzerland

Heptagenia sulphurea

Landa & Soldan 1985

distrib.

Heptagenia sulphurea

Hefti & Tomka & Zurwerra 1985

Heptagenia sulphurea

Braasch 1986

Mongolia

Heptagenia sulphurea

Herrera 1986

España

Heptagenia sulphurea

Herrmann & Andersson 1986

Heptagenia sulphurea

Otto & Sjostrom 1986

Heptagenia sulphurea

Tshernova & Kluge & 
Sinitshenkova & Belov 1986

distrib.

Heptagenia sulphurea

= Heptagenia dalecarlica

= Heptagenia soldatovi

Svensson 1986*

Sweden

Heptagenia sulphurea

Zurwera & Metzler 
& Tomka 1987*

biochem.

Heptagenia sulphurea

Grimm 1987* 

DeutschBR

Heptagenia sulphurea

Mol 1987*

Belgium

Heptagenia sulphurea

Boev & Bajanov 1987

s.Urals

Heptagenia sulphurea

Lehrke & Reusch 1987

Saxonia

Heptagenia sulphurea

Savolainen & Pulkkinen 1987*

Heptagenia sulphurea

Kluge 1987

Heptagenia (Heptagenia) sulphurea

= Heptagenia dalecarlica

= Heptagenia soldatovi

-"-

Heptagenia (Heptagenia) sulphurea f. sulphurea

-"-

Heptagenia (Heptagenia) sulphurea f. dalecarlica

-"-

Heptagenia (Heptagenia) sulphurea albicauda

Elliott & Humpesch & Macan 1988

Heptagenia sulphurea

Kluge 1988

Heptagenia (Heptagenia) sulphurea

Kluge 1989

, distrib.

Heptagenia (Heptagenia) sulphurea

-"-

, distrib.

Heptagenia (Heptagenia) sulphurea f. sulphurea

-"-

, distrib. 

Heptagenia (Heptagenia) sulphurea f. dalecarlica

-"-

, distrib. 

Heptagenia (Heptagenia) sulphurea albicauda

Landa & Soldan 1989*

Heptagenia sulphurea

Giller & Cambell 1989

substrat

Heptagenia sulphurea

Malzacher 1989

Heptagenia sulphurea

Bauernfeind 1990

Austria

Heptagenia sulphurea

Bazzanti 1991*

Heptagenia sulphurea

Krno 1991*

Slovakia

Heptagenia sulphurea

Savolainen & Hantula & 
Lokki & Saura 1991*

biochem.

Heptagenia sulphurea

≠ Heptagenia dalecarlica

Jazdzewska & Gorczynski 
1991:VI.Conf.Eph.

Heptagenia sulphurea

Bauernfeind 1992:93-97* 

Heptagenia sulphurea

D'Antonio 1992

Italia

Heptagenia sulphurea

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Heptagenia sulphurea

Devan 1993*

Heptagenia sulphurea

Devan 1993*

Heptagenia sulphurea

Csanyi 1994

Heptagenia sulphurea

Belfiore 1994*

Italia

Heptagenia sulphurea

Haybach & Fischer 1994*

Deutschland

Heptagenia sulphurea

Kluge & Silina 1994

Russia

Heptagenia sulphurea sulphurea

Huhta & Muotka & Tikkanen 1995

Heptagenia sulphurea

Sziraki 1995

Heptagenia sulphurea

Csanyi 1996

Heptagenia sulphurea

Thomas 1996*

s-e.France

Heptagenia sulphurea

Jacob & Dorn & Haybach 1996*

Heptagenia sulphurea

Novikova & Kluge 
1997:VIII.Conf.Eph.

W.Siberia

Heptagenia (Heptagenia) sulphurea f. sulphurea

Moog & al. 1997:VIII.Conf.Eph.

saprob.

Heptagenia sulphurea

Wise & O'Connor 1997:VIII.Conf.Eph.

Ireland

Heptagenia sulphurea

Csia & Sarkany-Kiss 1997

Romania

Heptagenia sulphurea

Parele & Ruzans & 
Spuris 1997*

Latvia

Heptagenia sulphurea

Kluge 1997:176-220

Heptagenia (Heptagenia) sulphurea   

-"-

Heptagenia (Heptagenia) sulphurea f. sulphurea

-"-

Heptagenia (Heptagenia) sulphurea f. dalecarlica

-"-

Heptagenia (Heptagenia) sulphurea albicauda

Juhasz & Turcsanyi & Kovacz & 
Olajos & Turcsanyi & Kiss1998

Heptagenia sulphurea

Sziraki 1998

Heptagenia sulphurea

Godunko 1998*

Ukraina

Heptagenia sulphurea

Galdean 1999

Heptagenia sulphurea

Weichselbaumer & 
Bauernfeind 1999*

Austria

Heptagenia sulphurea

Kovacs & Ambrus 
& Bankuti 1999*

Hungaria

Heptagenia sulphurea

Andrikovics & Kiss 2000

Heptagenia sulphurea

Haybach & Pottgiesser 2000*

N.Westfalen

Heptagenia sulphurea

Silina & Prokin 2000*

Lipetsk.prov.

Heptagenia sulphurea

Soldan & Puthz 2000*

chromosomes

Heptagenia sulphurea

Kelly-Quinn & Bracken 2000:1-223.

Ireland

Heptagenia sulphurea

Bauernfeind & Moog 2001

Heptagenia (Heptagenia) sulphurea

Andrikovics & Turcsanyi 2001

Heptagenia sulphurea

Kovacs 2001:87-92*

Hungaria

Heptagenia sulphurea

Kovacs & Ambrus 2001*

Hungaria

Heptagenia sulphurea

Kovacs & Juhasz 
& Turcsanyi 2001*

Hungaria

Heptagenia sulphurea

Kovacs & Ambrus 2002

Heptagenia sulphurea

Kovacs & Ambrus & 
Juhasz 2002:163-167*

Hungaria

Heptagenia sulphurea

Kovacs & Ambrus & 
Juhasz 2002:169-178*

Hungaria

Heptagenia sulphurea

Sziraki 2002

Heptagenia sulphurea

Kovacs & Ambrus & Juhasz 2003*

Hungaria

Heptagenia sulphurea

Kovacs & Bauernfeind 2003*

Hungaria

Heptagenia sulphurea

Beketov & Kluge 2003

W.Siberia

Heptagenia (Heptagenia) sulphurea

Arefina & Ivanov & Kocharina &
Lafer & Makarchenko & Teslenko
& Tiunova & Khamenkova 2003

Magadan.ob.

Heptagenia sulphurea

Haybach 2003:Lauterbornia

Heptagenia sulphurea

Wendling & Haybach 2003*

Romania

Heptagenia sulphurea

Kocharina & Khamenkova 2003

Magadan.ob.

Heptagenia sulphurea

Loskutova 2004

Polar Urals

Heptagenia sulphurea

Kluge 2004

thorax

in Heptagenia/f8=g7

= helvola [Ephemera]

= leucophthalma [Ephemera]

= ferruginea [Ephemera]

= costalis [Baetis]

= elegans [Baetis]

= straminea [Baetis]

= cyanops [Baetis]

= dalecarlica [Heptagenia]

= soldatovi [Heptagenia]

= albicauda [Heptagenia]

Wang & McCafferty 2004*

Heptagenia sulphurea

Tiunova & Potikha 2005

Far-East.Russia

Heptagenia sulphurea

Reznic 2005

Yakutia

Heptagenia sulphurea

Leman & Esin & Chalov
& Chebanova 2005

Kamchatka

Heptagenia sulphurea

Bauernfeind & Kovacs 
& Ambrus 2005*

Hungaria

Heptagenia sulphurea

Kovacs 2005:101-110*

Hungaria

Heptagenia sulphurea

Kovacs 2006:143-158*

Hungaria

Heptagenia sulphurea

Kovacs 2006:325-331*

Hungaria

Heptagenia sulphurea

Kovacs & Kovacs 2006:159-165*

Hungaria

Heptagenia sulphurea

Peeters & Brugmans 
& Beijer & Fraken 2006

Heptagenia sulphurea

Prokopov  & Godunko 2007

Heptagenia sulphurea

sensu Kiseleva & Ezernitsky 1985 
?= Heptagenia samochai 

Tiunov & Tiunva 2007*

Heptagenia sulphurea

Sharapova 2007

W.Siberia

Heptagenia sulphurea

Enkhtaivan & Soldan 2008:XI.Conf.Eph.*

Mongolia

Heptagenia sulphurea

Beketov 2008:289-299

Heptagenia sulphurea

Tiunova 2008:172-188

Amur.prov.

Heptagenia sulphurea

Kovacs & Ambrus & Juhasz 
& Olajos & Szilagyi 2008 '

Lithuania

Heptagenia sulphurea

Tiunova & Teslenko & Reznik 2009

Yakutia

Heptagenia sulphurea

Savolainen 2009:1-35

Finland

Heptagenia sulphurea

Kovacs 2009:73-85*

Hungaria

Heptagenia sulphurea

Sroka 2009:XII.Conf.Eph.

mola; food

Heptagenia sulphurea

Soldan & Enktaivan & 
Godunko 2009:XII.Conf.Eph.

Mongolia

Heptagenia sulphurea

= cyanops [Baetis]

?= soldatovi [Heptagenia]

Tiunova 2009:XII.Conf.Eph.

Far-East.Russia

Heptagenia sulphurea

Kluge 2009:109-134

E.Siberia,
Mongolia

in Heptagenia/f8=g7

Tiunova & Bazova 2010

r.Selenga

Heptagenia (Heptagenia) sulphurea

Kjaerstad & Webb & Ekrem 2012

N.Norwegia

Heptagenia sulphurea

Bauernfeind & Soldán 2012

Heptagenia (s.str.) sulphurea

Potikha 2015

Far-East.Russia

Heptagenia sulphurea

Erdenee & Maasri & Gelhaus & Bayartogtokh 2016

Mongolia

Heptagenia sulphurea

Bauernfeind 2018

Albania

Heptagenia (Heptagenia) sulphurea

Evseeva 2021

Kazakhstan: bass.Irtysh

Heptagenia sulphurea

Kluge 2022:151-186.

tarsus

Heptagenia sulphurea

Mangot 2023

France

Heptagenia sulphurea

Heptagenia sulphurea

Heptagenia (Heptagenia) sulphurea f. sulphurea

--/

-

in Heptagenia/f8=g7

= helvola [Ephemera]

= leucophthalma [Ephemera]

= ferruginea [Ephemera]

= costalis [Baetis]

= elegans [Baetis]

= straminea [Baetis]

= cyanops [Baetis]

= dalecarlica [Heptagenia]

= soldatovi [Heptagenia]

= albicauda [Heptagenia]