EPHEMEROPTERA OF THE WORLD

arb.sig. ABC

z   

Heptagenia/fg 

f: Heptageniinae Needham & Betten 1901: 419

g: Heptagenia Walsh 1863: 197, typus Palingenia flavescens Walsh 1863 (design. Eaton 1868b: 90)

Nomina circumscribentia:


Heptagenioidea

Heptageniidae

Heptageniinae

Heptageniini

gen. Heptagenia

subgen. Heptagenia

taxa subordinated to Heptagenia/g1 & Heptagenioidea & Heptageniidae & Heptageniinae & Heptagenia

binomina

Heptageniidae genn.spp.

Heptagenia spp.


Heptagenia/f1

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 248

 

Heptagenia/f1=Oligoneuria/g1 
Branchitergaliae  

 

Bidentiseta
= Ephemera/fg5

Eusetisura

Heptagennota

Kluge 2004 

Heptagenia/f1=Oligoneuria/g1 
= Branchitergaliae

= Heptagenioidea: 
McCafferty 1997

Bidentiseta
= Ephemera/fg5

Eusetisura

Heptagennota

 

 

 

 

 

 

 

Heptagenia/f1=Oligoneuria/g1  
= Branchitergaliae

 

Bidentiseta

Eusetisura

Heptagennota

 

 

 

 

 

 


Heptagenia/f2

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 248

 

Heptagenia/f2=g1 = Heptagennota

 

Branchitergaliae

Pseudron

Pentamerotarsata

Kluge 2004: 158

Heptagenia/f2=g1 = Heptagennota

= Heptageniidae: 
Edmunds & Traver 1954

= Heptagenioidea: 
Kluge & al. 1995

Branchitergaliae

Pseudron

Pentamerotarsata

 

 

 

 

 

 

 

Heptagenia/f2=g1 = Heptagennota

 

Branchitergaliae

Pseudron

Pentamerotarsata

 

 

 

 

 

 


Heptagenia/f3

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 251

 

Heptagenia/f3=g2 = Pentamerotarsata

 

Heptagennota

Arthroplea

Radulapalpata

Kluge 2004

Heptagenia/f3=g2 = Pentamerotarsata

= Heptageniinae: 
Needham 1901

= Heptageniidae: 
Lestage 1917

Heptagennota

Arthroplea

Radulapalpata

 

 

 

 

 

 

 

Heptagenia/f3=g2 = Pentamerotarsata

 

Heptagennota

Arthroplea

Radulapalpata

 

 

 

 

 

 


Heptagenia/f4

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 251

 

Heptagenia/f4=g3 = Radulapalpata

 

Pentamerotarsata

 

Kluge 2004: 165 

Heptagenia/f4=g3 = Radulapalpata

= Ecdyonuridae: Balthasar 1937

= Heptageniidae: Landa 1969

= Heptageniinae: Kluge 1988, 1993

Pentamerotarsata

Heptagenia/f5=g4

Raptoheptagenia

Cinygma

Rhithrogena/fg1

 

 

 

 

 

 

 

 

Heptagenia/f4=g3 = Radulapalpata

 

Pentamerotarsata

Heptagenia/f5=g4

Raptoheptagenia

Cinygma

Rhithrogena/fg1

 

 

 

 

 

 


Heptagenia/f5

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 166

Heptagenia/f5=g4

=  Heptageniini: 
Kluge 1988

Radulapalpata

Heptagenia/f6=g5

Ecdyonurus/fg1

 

 

 

 

 

 

 

Heptagenia/f5=g4

 

Radulapalpata

Heptagenia/f6=g5

Ecdyonurus/fg1

 

 

 

 

 

 


Heptagenia/f6

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 168

Heptagenia/f6=g5

= gen. Heptagenia
Kluge 1988

Heptagenia/f5=g4

Kageronia 

Heptagenia/f7=g6

 

 

 

 

 

 

 

Heptagenia/f6=g5

 

Heptagenia/f5=g4

Kageronia 

Heptagenia/f7=g6

 

 

 

 

 

 


Heptagenia/f7

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2002: 169

Heptagenia/f7=g6

 

Heptagenia/f6=g5

Heptagenia/f8=g7

Dacnogenia 

Stenonema/fg1

Kluge 2022:151-186. tarsus Heptagenia/f7=g6   Heptagenia/f6=g5  

 

 

 

 

 

 

 

Heptagenia/f7=g6

 

Heptagenia/f6=g5

Heptagenia/f8=g7

Dacnogenia 

Stenonema/fg1

 

 

 

 

 

 


Heptagenia/f8

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 169

Heptagenia/f8=g7

= subgen. Heptagenia: Kluge 1988

Heptagenia/f7=g6

 

 

 

 

 

 

 

 

Heptagenia/f8=g7

 

Heptagenia/f7=g6

 

 

 

 

 

 

 


superfamilia & subordo Heptagenioidea

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Ulmer 1920:97–144*

Heptagenioidea (subordo)

 

Ephemeroptera

Baetiscidae

Siphlonuridae

Ametropodidae

Ecdyonuridae = Heptageniidae

Lestage 1924

 

Heptagenioidea (subordo)

 

 

Ecdyonuridae

Ulmer 1929

 

Heptagenioidea (subordo)

 

 

Ecdyonuridae

Edmunds & Traver 1954

 

Heptagenioidea

= Siphlonuroidea

= Baetoidea

 

Siphlonuridae

Isonychiidae

Oligoneuriidae

Ametropodidae

Baetidae

Siphlaenigmatidae

Heptageniidae

Edmunds & Allen 1957*

 

Heptagenioidea

 

Ephemeroptera

(-"-: Edmunds & Traver 1954)

Demoulin 1958

 

Heptagenioidea

 

 

Ametropodidae

Hetageniidae

Leptophlebiidae

Tshernova 1962

 

Heptageniidea

 

Euplectoptera

Ametropodide

Heptageniidae

Edmunds 1962

 

Heptagenioidea

 

 

Siphlonuridae

Oligoneuriidae

Ametropodidae

Baetidae

Heptageniidae

Edmunds & Allen & Peters 1963

 

Heptagenioidea

 

 

Siphlonuridae

Oligoneuriidae

Ametropodidae

Baetidae

Siphlaengmatidae

Heptageniidae

Tshernova 1970

 

Heptagenioidea

 

 

Heptageniidae

†Epeoromimidae

Riek 1970

 

Heptagenioidea

 

 

Heptageniidae

Riek 1973

 

Heptagenioidea

 

 

Heptageniidae

Koss 1973

 

Heptagenioidea

 

 

Siphlonuridae

Oligoneuriidae

Ametropodidae

Metretopodidae

Baetidae

Siphlaengmatidae

Heptageniidae

Edmunds & Jensen & Berner 1976

 

Heptagenioidea

 

 

Siphlonuridae

Metretopodidae

Siphlaenigmatidae

Baetidae

Oligoneuriidae

Heptageniidae

Ametropodidae

McCafferty & Edmunds 1979

 

syn. Baetoidea

 

 

 

Tshernova 1980

 

Heptagenioidea

 

 

Heptageniidae

†Epeoromimidae

Landa & Soldan 1985

 

Heptagenioidea

 

Schistonota

†Epeoromimidae

Oligoneuriidae

Heptageniidae

McCafferty 1990

phylog.

Heptagenioidea

 

 

Isonychiidae

Coloburiscidae

Oligoneuriidae

Heptageniidae

Peters & Campbell 1991

 

Heptagenioidea

 

Schistonota

Isonychiidae

Coloburiscidae

Oligoneuriidae

Heptageniidae

Kluge & Studemann & Landolt & Gonser 1995: 105

 

Heptagenioidea

 

Setisura

 

Wang & McCafferty 1995

 

Heptagenioidea

 

 

Heptageniidae

Pseudironidae

Arthropleidae

McCafferty 1997: 96

 

Heptagenioidea

 

 

 

Kluge 1997:176-220

 

Heptagenioidea

 

 

Heptageniidae

Arthropleidae

 

 

Heptagenioidea

 

 

 

 

 

Heptagenia/f1 
Heptagenia/f2=g1

Heptagenia/f3=g2

Heptagenia/f4=g3

 

 

 

 

 

 

 

 

 


familia Heptageniidae

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Lestage 1917

Heptageniidae

 

 

Rhithrogeninae

Heptageniinae

Bengtsson 1917

 

Heptageniidae

 

 

 

Spieth 1933

phylog.

Heptageniidae

 

 

 

Needham & Traver & Hsu 1935

Heptageniidae

 

 

Heptageniinae

Edmunds & Traver 1954a: 237

Heptageniidae

 

Heptagenioidea

Heptageniinae

Pseudironinae

Demoulin 1958

 

Heptageniidae

 

Heptagenioidea

 

Demoulin 1961

 

Heptageniidae

 

Heptagenioidea

 

Edmunds & Traver 1959

 

Heptageniidae

 

 

Heptageniinae

Anepeorinae

Tshernova 1962*

Heptageniidae

 

Heptagenioidea

 

Edmunds & Allen & Peters 1963

 

Heptageniidae

 

Heptagenioidea

Heptageniinae

Anepeorinae

Pseudironinae

Arthropleinae

Landa 1969a: 153

Heptageniidae

 

 

 

Riek 1970

 

Heptageniidae

 

Heptagenioidea

 

Tshernova 1970

 

Heptageniidae

 

Heptagenioidea

Heptageniinae

Riek 1973

 

Heptageniidae

 

Heptagenioidea

 

Koss 1973

 

Heptageniidae

 

Heptagenioidea

 

Landa 1973

Heptageniidae

 

 

(excl. Arthropleidae)

Edmunds & Jensen & Berner 1976

Heptageniidae

 

Heptagenioidea

Pseudironinae

Heptageniinae

Arthropleinae

Anepeorinae

Spinadinae

McCafferty & Edmunds 1979

 

Heptageniidae

 

Baetoidea

Heptageniinae

Anepeorinae

Pseudironinae

Spinadinae

Arthropleinae

Tshernova 1980

 

Heptageniidae

 

Heptagenioidea

 

Edmunds 1982*

Heptageniidae

 

Baetoidea

 

Landa & Soldan 1985

 

Heptageniidae

 

Heptagenioidea

Arthropleinae

Heptageniinae

Anepeorinae

Spinadinae

Tomka & Zurwerra 1985

 

Heptageniidae

 

 

Heptageniinae

Pseudironinae

Arthropleinae

Tshernova & Kluge & 
Sinitshenkova & Belov 1986

Heptageniidae

 

 

 

Hubbard 1988

 

Heptageniidae

 

Baetoidea

Heptageniinae

Arthropleinae

Kluge 1989

 

Heptageniidae

 

Costatergalia = Baetoidea

 

McCafferty 1990

 

Heptageniidae

 

Heptagenioidea

 

Tomka & Elpers 1991:VI.Conf.Eph.*

 

Heptageniidae

 

Ephemeroptera

Heptageniinae

Anepeorinae

Spinadinae

McCafferty 1991

 

Heptageniidae

 

Setisura

 

McCafferty 1991

phylog.pos.

Heptageniidae

= Arthropleidae

 

 

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Heptageniidae

 

Ephemeroptera

 

Wang & McCafferty 1995

 

Heptageniidae

 

Heptagenioidea

 

Kluge 1997:176-220

 

Heptageniidae

 

Heptagenioidea

 

Wang & McCafferty 2004

 

Heptageniidae

 

 

Ecdyonurinae

Heptageniinae

Rhithrogeninae

 

 

Heptageniidae

 

 

 

 

 

Heptagenia/f1 
Heptagenia/f2=g1

Heptagenia/f3=g2

Heptagenia/f4=g3

 

 

 

 

 

 

 

 

 


subfamilia Heptageniinae

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Needham & Betten 1901:419*

Heptageniinae subfam.n. = Heptagenia s.l.

 

Ephemeridae

-

Needham 1905*

Heptageniinae

 

Ephemeridae

Epeorus

Iron

Ecdyurus

Cinygma

Rhithrogena

Heptagenia

et al.

Lameere 1917

 

Heptageniinae

 

Ephemeridae

Heptageniini

Ametropini

Lestage 1917

Heptageniinae

 

Heptageniidae

Heptagenia

Ecdyurus

Arthroplea

Epeorus

Cnygma

Needham & Traver & Hsu 1935

 

Heptageniinae

 

Heptageniidae

Cinygma

Cinygmula

Heptagenia

Iron

Ironodes

Ironopsis

Rhithrogena

Stenonema

Anepeorus

Arthroplea

Edmunds & Traver 1954

 

Heptageniinae

 

Heptageniidae

Afronurus

Atopopus

Bleptus

Cinygma

Cinygmina

Cinygmula

Compsoneuria

Compsoneuriella

Ecdyonurus

Epeorella

Epeorus

Heptagenia

Notonurus

Ororotsia

Paegniodes

Rhithrogena

Rhithrogenella

Stenonema

Thalerosphyrus

Arthroplea

Anepeorus

Edmunds & Allen & Peters 1963

 

Heptageniinae

 

Heptageniidae

Afronurus

Atopopus

Bleptus

Cinygma

Cinygmina

Cinygmula

Compsoneuria

Compsoneuriella

Ecdyonurus

Epeorella

Epeorus

Heptagenia

Notonurus

Ororotsia

Paegniodes

Rhithrogena

Rhithrogenella

Stenonema

Thalerosphyrus

Tshernova 1970*

 

Heptageniinae

 

Heptageniidae

Afronurus

Afghanurus

Atopopus

Bleptus

Cinygma

Cinygmina

Cinygmula

Compsoneuria

Compsoneuriella

Ecdyonurus

Epeiron

Epeorella

Epeorus

Heptagenia

Notonurus

Ororotsia

Paegniodes

Rhithrogena

Rhithrogenella

Sigmoneuria

Stenonema

Thalerosphyrus

Succinogenia

Miocenogenia

Edmunds & Jensen & Berner 1976

Heptageniinae

 

Heptageniidae

 

McCafferty & Edmunds 1979

 

Heptageniinae

 

Heptageniidae

 

Tomka & Zurwerra 1985

 

Heptageniinae

 

Heptageniidae

Afronurus

Atopopus

Bleptus

Cinygma

Cinygmina

Cinygmula

Compsoneuria

Compsoneuriella

Ecdyonuroides

Ecdyonurus

Electrogena

Epeorella

Epeorus

Heptagenia

Iron

Ironodes

Ironopsis

Nixe

Leucrocuta

Macdunnoa

Notacanthurus

Paegniodes

Rhithrogena

Rhithrogenella

Sigmoneuria

Stenacron

Stenonema

Thalerosphyrus

Anepeorus

Spinadis

Landa & Soldan 1985

 

Heptageniinae

 

Heptageniidae

32 genn.

Kluge 1988

 

Heptageniinae

 

Heptageniidae

Heptagenini

Cinygmini

Rhithrogenini

Tomka & Elpers 1991

phylog.

Heptageniinae

 

Heptageniidae

 

Kluge 1993a

 

Heptageniinae

 

 

 

Wang & McCafferty 2004

 

Heptageniinae

 

Heptageniidae

Compsoneuriini

Heptageniini

Kageroniini

Stenonematini

Webb & McCafferty 2008

Heptageniinae

 

Heptageniidae

Dacnogenia

Heptagenia

Kageronia

Maccaffertium

Macdunnoa

Raptoheptagenia

Stenacron

Stenonema

Trichogenia

 

 

Heptageniinae

 

Heptageniidae

 

 

 

Heptagenia/f1 
Heptagenia/f2=g1

Heptagenia/f3=g2

Heptagenia/f4=g3

Heptagenia/f5=g4

Heptagenia/f6=g5

Heptagenia/f7=g6

Heptagenia/f8=g7

 

 

 

 

 

 

 

 

 


tribus Heptageniini

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Lameere 1917

 

Heptageniini

 

Ephemeridae-Heptageninae

Heptagenia

Ecdyonurus

Rhithrogena

Epeorus

Iron

Kluge 1988: 292

 

Heptageniini

 

Heptagenidae

Heptagenia s.l.

Ecdyonurus s.l.

Wang & McCafferty 2004

 

Heptageniini

 

Heptageniinae

Heptagenia

Dacnogenia

Raptoheptagenia

 

 

Heptageniini

 

 

 

 

 

Heptagenia/f1 
Heptagenia/f2=g1

Heptagenia/f3=g2

Heptagenia/f4=g3

Heptagenia/f5=g4

Heptagenia/f6=g5

Heptagenia/f7=g6

Heptagenia/f8=g7

 

 

 

 

 

 

 

 

 


genus Heptagenia

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Walsh 1863:167–272:Note16*

Heptagenia gen.n.

= Palingenia subgen.B: Walsh 1862

= diff.6: Hagen in Walsh 1863

Ephemerina

 

Eaton 1868:82-91*

Heptagenia

= Ecdyurus syn.n.

Ephemeridae

[sect.A]

sect.B,venosa

Eaton 1871*

Heptagenia

= Ecdyurus

Ephemeridae

 

Eaton 1881:(18)21-27*

Heptagenia

Ecdyurus

Ephemeridae

 

Eaton 1883-1888

Heptagenia restr.Etn.1881

 

Ephemeridae

section 14 of Ecdyurus

 

Schiller 1890

Heptagenia

 

 

 

Banks 1900: 246

 

Heptagenia

 

Ephemeridae-Leptophlebini tr.n.

 

Needham 1905*

Heptagenia

 

Ephemeridae-Heptageniinae

 

Jacobson & Bianchi 1905

 

Heptagenia

 

Siphlonuridae-Ecdyurinae

 

Banks 1907:1-53

ref.

Heptagenia

= Ecdyonurus

= Ecdyurus

= Rhithrogena syn.n.

= Iron syn.n.

= Cinygma syn.n.

 

 

Lameere 1917

 

Heptagenia

 

Heptageniini

 

Lestage 1917

Heptagenia

 

Heptageniidae-Heptageniinae

 

Lestage 1919

Heptagenia

 

Heptageniinae

 

Ulmer 1920:97–144*

Heptagenia

 

Ecdyonuridae = Heptageniidae

 

McDunnough 1924

 

Heptagenia

 

 

 

Scheoenemund 1930

Heptagenia

 

 

 

Traver 1932:85-161

Heptagenia

! Traver 1933

 

 

Traver 1933:105-125*

Heptagenia

= Ecdyonurus sensu Traver 1932

sensu Traver 1932
= Stenonema

Heptageniinae

 

Traver 1933

 

Heptagenia

 

 

 

Spieth 1933

phylog.

Heptagenia

 

Heptageniidae

 

Needham & Traver & Hsu 1935

Heptagenia

= Ecdyurus: Needham 1905

Heptageniidae-Heptageniinae

 

Imanishi 1935

 

Heptagenia

= Kageronia syn.n.

 

 

Ulmer 1939*

Heptagenia

 

Ecdyonuridae

 

Burks 1953

Heptagenia

 

Heptageniidae

gr.persimplex

gr.elegantula

gr.pulla

gr.flavescens

gr.lucidipennis

gr.maculipennis

Edmunds & Traver 1954

 

Heptagenia

 

Heptageniidae-Heptageniinae

 

Bogoescu & Tabacaru 1962

Heptagenia

 

 

 

Edmunds & Allen & Peters 1963

 

Heptagenia

 

Heptageniidae-Heptageniinae

 

Tshernova 1970*

 

Heptagenia

 

Heptageniidae-Heptageniinae

 

Edmunds & Jensen & Berner 1976

Heptagenia

 

Heptageniidae-Heptageniinae

 

Kluge 1980

Heptagenia

 

Heptageniidae

 

Yoon & Bae 1984*

Heptagenia

 

 

 

Tomka & Zurwerra 1985

Heptagenia

 

Heptageniidae-Heptageniinae

 

Kluge 1983b: 16 (Abstract)

 

Heptagenia

[= Sigmoneuria syn.n.]

[= Stenonema syn.n.]

[= Stenacon syn.n.]

[= Madunoa syn.n.]

Heptageniidae

subg.Heptagenia

subg.Dacnogenia

subg.Parastenacron

subg.Stenacon

subg.Stenonema

subg.Macdunoa

Tshernova & Kluge & 
Sinitshenkova & Belov 1986

Heptagenia

 

Heptageniidae

 

Braasch 1986*

Heptagenia

 

 

 

Kluge 1987

Heptagenia

= Sigmoneuria syn.n.

 

 

Kluge 1988: 292

Heptagenia

= Sigmoneuria 

= Stenonema syn.n.

= Stenacon syn.n.

= Macdunoa syn.n.

Heptageniini

subg.Kageronia

subg.Dacnogenia

subg.Heptagenia

subg.Stenonema

subg.Stenacon

subg.Macdunoa

Hefti & Tomka 1985

Heptagenia

 

Heptageniidae-Heptageniinae

 

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Heptagenia

 

Heptageniidae

 

Jacob & Dorn & Haybach 1996

Heptagenia

 

 

 

Kluge 1997:176-220

(in Russia)

Heptagenia s.l.

 

Heptageniidae

subg.Kageronia

subg.Dacnogenia

subg.Heptagenia

et al.

Wang & McCafferty 2004:44

 

Heptagenia

 

Heptageniinae-Heptageniini

 

Haybach 2006:39-44'

Heptagenia

 

Heptagheniidae

 

Webb & McCafferty 2008

Heptagenia

 

Heptageniinae

 

Burian 2019

Heptagenia

 

 

 

 

Heptagenia

 

 

 

 

 

Heptagenia/f1 
Heptagenia/f2=g1

Heptagenia/f3=g2

Heptagenia/f4=g3

Heptagenia/f5=g4

Heptagenia/f6=g5

Heptagenia/f7=g6

Heptagenia/f8=g7

 

 

 

 

 

 

 

 

 


subgenus Heptagenia

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 1987

subgen. Heptagenia

= Sigmoneuria syn.n.

Heptagenia

 

Kluge 1988: 295

subgen. Heptagenia

= Sigmoneuria

Heptagenia

 

Kluge 1997:176-220

 

subgen. Heptagenia

= Sigmoneuria

Heptagenia 

 

 

 

subgen. Heptagenia

 

 

 

 

 

Heptagenia/f7=g6

Heptagenia/f8=g7

 

 

 

 

 

 

 

 

 


taxa subordinated to Heptagenia/g1 & Heptagenioidea & Heptageniidae & Heptageniinae & Heptagenia


binomina:

Heptagenia

 abnormis Tshernova 1949 [Heptagenia] — in Cinygma/fg1

adaequata McDunnough 1924 [Heptagenia]

affine Traver 1933 [Stenonema]

 affinis Eaton 1885 [Ecdyurus] — in Ecdyonurus/fg1

alabamae Traver 1937 [Stenonema]

albicauda Kluge 1987 [Heptagenia sulphurea]

allegheniense Carle 1977 [Stenonema]

alpicola Eaton 1871 [Heptagenia] — typus nominis Alpiron — in Epeorus/fg3

amseli Demoulin 1964 [Sigmoneuria]  

angustipennis Rambur 1842 [Ephemera]

annexum Traver 1933 [Stenonema]  

annulifera Walker 1860 [Palingenia]

aphrodite McDunnough 1926 [Heptagenia] — in Ecdyonurus/fg1

areion Burks 1953 [Stenonema]

ares Burks 1953 [Stenonema]

arsenjevi Tshernova 1952 [Heptagenia]

arwini Bednarik & McCafferty 1979 [Stenonema mediopunctatum]

 assimilis Eaton 1885 [Epeorus] — in Epeorus/fg1

atypica Demoulin 1968 [Heptagenia]

aurantiaca Burmeister 1839 [Baetis] — in Rhithrogena/fg3

bachofeni Demoulin 1968 [Heptagenia]

basalis Walker 1853 [Baetis] — in Metretopus/fg1

bednariki McCafferty 1981 [Stenonema]

bella Allen & Cohen 1977 [Heptagenia] — in Ecdyonurus/fg1

 bellieri Hagen 1860 [Baetis] — in Ecdyonurus/fg1

bellum Traver 1933 [Stenonema]  

bipunctata Esben-Petersen 1916 [Heptagenia]  

bipunctatus McDunnough 1926 [Ecdyonurus]  

birdi Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

borealis Eaton 1871 [Heptagenia] — in Metretopus/fg1

brunnea Hagen 1875 [Heptagenia] — in Rhithrogena/fg3

brunnea Flowers 1982 [Macdunnoa]

californicus Banks 1910 [Heptagenia (Epeorus)] — in Epeorus/fg2-Ironodes

canadensis Walker 1853 [Baetis]  

candidum Traver (in Needham & Traver & Hsu) 1935 [Stenonema]

carlsoni Lewis 1974 [Stenonema]

carolina Banks 1914 [Heptagenia]

cerea Pictet 1843 [Baetis]  

chinensis Ulmer 1920 [Heptagenia]

citrina Hummel 1825 [Ephemera] nomen oblitum  

 coerulans Rostock 1878 [Heptagenia]

 concii Grandi 1953 [Heptagenia]   — in Ecdyonurus/fg1

confinis Tshernova 1928 [Ecdyonurus]  

conjunctum Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

convergens Lestage 1924 [Ecdyonurus]  

costalis Curtis 1834 [Baetis]  

costata Navás 1936 [Heptagenia]

couzonica Real 1947 [Heptagenia sulphurea]  

coxalis Banks 1914 [Heptagenia]  

criddlei McDunnough 1927 [Heptagenia] — in Ecdyonurus/fg1

cruentata Walsh 1863 [Heptagenia] — typus nominis Raptoheptagenia  

culacantha Evans & Botts & Flowers 1985 [Heptagenia]

cupulata Eaton 1871 [Heptagenia] — typus nominis Paegniodes — in Rhithrogena/fg1

cyanops Pictet 1843 [Baetis]  

dalecarlica Bengtsson 1912 [Heptagenia]  

 davidica Navás 1935 [Heptagenia]

denervosa Tiensuu 1935 [Heptagenia sulphurea]  

dentata Braasch 1979 [Heptagenia] — in Ecdyonurus/fg1

 determinata Walker 1853 [Baetis] — typus nominis Thalerosphyrus — in Ecdyonurus/fg1

diabasia Burks 1947 [Heptagenia]

dolosa Traver (in Needham & Traver & Hsu) 1935 [Heptagenia]

elegans Curtis 1834 [Baetis]  

elegantula Eaton 1885 [Rhithrogena]

exiguum Traver 1933 [Stenonema]

 fallax Hagen 1864 [Baetis] — in Ecdyonurus/fg1

femorata Say 1823 [Baetis]

ferruginea Gmelin 1790 [Ephemera]  

flava Rostock 1878 [Heptagenia]

flavata Navás 1922 [Heptagenia]

flaveola Pictet 1843 [Baetis]

flavescens Walsh 1862 [Palingenia] — typus nominis Heptagenia

flavianula McDunnough 1924 [Heptagenia] — in Rhithrogena/fg3

flavipennis Dufour 1841 [Ephemera]  

floridense Lewis 1974 [Stenonema]

fluminea Zhang 1989 [Heptagenia]

 fluminum Pictet 1843 [Baetis] — in Ecdyonurus/fg1

 forcipula Pictet 1843 [Baetis] — in Ecdyonurus/fg1

frontale Banks 1910 [Heptagenia]  

fusca Walker 1853 [Baetis] nom.praeocc. — syn.obj. jejuna [Rhithrogena]

fusca Clemens 1913 [Heptagenia]  

fuscogrisea Retzius 1783 [Ephemera]

gallica Eaton 1885 [Heptagenia]  

geminatum Eaton 1885 [Cinygma] — in Epeorus/fg2-Ironodes/g1

gemmata Scopoli 1763 [Ephemera]

germanica Eaton 1885 [Rhithrogena] — in Rhithrogena/fg3

gildersleevei Traver (in Needham & Traver & Hsu) 1935 [Stenonema]

gleissi Demoulin 1968 [Heptagenia]

 gridellii Grandi 1954 [Heptagenia] — in Ecdyonurus/fg1

guranica Belov 1981 [Heptagenia]

 guttata Pictet 1843 [Baetis]

hazaraensis Ali 1970 [Heptagenia]

hebe McDunnough 1924 [Heptagenia] — in Ecdyonurus/fg1

helvola Subz. 1776 [Ephemera]  

heterotarsalis McDunnough 1933 [Ecdyonurus]  

horrida McDunnough 1926 [Heptagenia] — in Ecdyonurus/fg1

impersonata McDunnough 1925 [Heptagenia] — in Rhithrogena/fg3

inconspicua McDunnough 1924 [Heptagenia] — in Ecdyonurus/fg1

 insignis Eaton 1870 [Heptagenia] — in Ecdyonurus/fg1

integer McDunnough 1924 [Heptagenia]  

 integrum Eaton 1885 [Cinygma] — typus nominis Cinygma

interpunctata Say 1839 [Baetis]

iridina Kolenati 1859 [Baetis] — in Rhithrogena/fg3

ithaca Clemens & Leonard 1924 [Heptagenia]

jejuna Eaton 1885 [Rhithrogena] — in Rhithrogena/fg3

jewetti Allen 1966 [Heptagenia] — in Ecdyonurus/fg1

joernensis Bengtsson 1909 [Ecdyurus] — in Ecdyonurus/fg1

julia Traver 1933 [Heptagenia]

juno McDunnough 1924 [Heptagenia] — in Ecdyonurus/fg1

kennedyi McDunnough 1924 [Heptagenia] — in Ecdyonurus/fg1

kibunensis Imanishi 1936 [Ecdyonurus] — in Ecdyonurus/fg1

kihada Matsumura 1931 [Heptagenia]

kyotoensis Gose 1963 [Heptagenia]

lateralis Curtis 1834 [Baetis] — typus nominis Electrogena — in Ecdyonurus/fg1

lenati McCafferty 1990 [Stenonema]

lepton Burks 1947 [Stenonema]  

leucophthalma Ström 1783 [Ephemera]  

lieftincki Ulmer 1939 [Heptagenia]

ligata Demoulin 1968 [Heptagenia]

limbata Navás 1936 [Heptagenia]

longicauda Stephens 1835 [Baetis]

longimanus Eaton 1883 [Epeorus] — typus nominis Iron

lucidipennis Clemens 1913 [Ecdyurus] — typus nominis Nixe

luridipennis Burmeister 1839 [Baetis] — in Siphlonurus/fg1

lutea Clemens 1913 [Heptagenia]

lutea Kluge 1987 [Heptagenia (Heptagenia)] nom. praeocc. — syn.subj. samochai [Sigmoneuria]

 macedonica Ikonomov 1954 [Heptagenia] — in Ecdyonurus/fg1

maculata Poda 1761 [Ephemera] — in Ecdyonurus/fg1

maculipennis Walsh 1863 [Heptagenia] typus nominis Leucrocuta

majus Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

manifesta Eaton 1885 [Rhithrogena] — in Rhithrogena/fg3

marginalis Banks 1910 [Heptagenia]

maxillaris Braasch & Soldán 1988 [Trichogenia]

 mazedonica Ikonomov 1954 [Heptagenia] — in Ecdyonurus/fg1

mediopunctatus McDunnough 1926 [Ecdyonurus]

meririvulanum Carle & Lewis 1978 [Stenonema]

metriotes Burks 1953 [Stenonema]  

mexicana Ulmer 1920 [Heptagenia]

 micracantha Kluge 1989 [Heptagenia (Dacnogenia) coerulans  

mimus Eaton 1885 [Cinygma] — in Cinygmula/g1

minerva McDunnough 1924 [Heptagenia] — in Ecdyonurus/fg1

minnetonka Daggy 1945 [Stenonema]

minor You & Gui 1995 [Heptagenia]

modestus Banks 1910 [Heptagenia (Epeorus)]

molinai Navás 1930 [Heptagenia] — typus nominis Dictyosiphlon — in Eusetisura

mongolica Bajkova & Varykhanova 1978 [Heptagenia] — in Ecdyonurus/fg1

montana Pictet 1843 [Baetis]

morrisoni Banks 1924 [Epeorus] — in Rhithrogena/fg3

nasuta Ulmer 1939 [Heptagenia]

nepotellus McDunnough 1933 [Ecdyonurus]  

ngi Hsu 1936 [Heptagenia]

nigrescens Klapálek (in Dziedzielewicz & Klapálek) 1908 [Heptagenia] — in Ecdyonurus/fg1

nigrimana Dufour 1841 [Ephemera] — in Ecdyonurus/fg1

nipawinia Lehmkuhl 1979 [Macdunnoa]

nitidus Eaton 1885 [Iron] — typus nominis Ironodes

nivata Eaton 1871 [Heptagenia] — in Rhithrogena/fg3

nubila Kimmins 1937 [Heptagenia]

obscura Stephens 1835 [Baetis] — in Ephemerella/fg1

ohioense Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

orbiticola Kluge 1987 [Heptagenia (Kageronia)]

otiosa McDunnough 1935 [Heptagenia] — in Ecdyonurus/fg1

 ozrensis Tanasijevic 1975 [Heptagenia] — in Ecdyonurus/fg1

pallidum Traver 1933 [Stenonema]

 par Eaton 1885 [Cinygma] — in Cinygmula/g1

patoka Burks 1947 [Heptagenia]

pectoralis Matsumura 1931 [Heptagenia]

perfida McDunnough 1926 [Heptagenia] — in Ecdyonurus/fg1

perflava Brodsky 1930 [Heptagenia]

persimplex McDunnough 1929 [Heptagenia]

petersi Allen 1966 [Heptagenia] — in Ecdyonurus/fg1

picteti Meyer-Dür 1864 [Baetis] — in Ecdyonurus/fg1

placita Banks 1910 [Heptagenia]  

platysma Tshernova 1941 [Heptagenia]  

pleuralis Banks 1910 [Heptagenia (Epeorus)] — in Epeorus/fg1

proximum Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

pudica Hagen 1861 [Ephemera]

pulchella Walsh 1862 [Palingenia]

pullus Clemens 1913 [Ecdyurus]

purpurascens Pictet 1843 [Baetis] — in Ecdyonurus/fg1

 quadrilineata Landa 1969 [Heptagenia] — in Ecdyonurus/fg1

quadripunctata Kluge 1989 [Heptagenia (Heptagenia)]

quebecensis Provancher 1876 [Cloe] — in Leptophlebia/fg1

querula McDunnough 1924 [Heptagenia]  

quinquespinum Lewis 1974 [Stenonema]  

reversalis McDunnough 1924 [Heptagenia]

rivulicolus McDunnough 1933 [Ecdyonurus]  

rodocki Traver (in Needham & Traver & Hsu) 1935 [Heptagenia] — in Ecdyonurus/fg1

rosea Traver (in Needham & Traver & Hsu) 1935 [Heptagenia] — in Ecdyonurus/fg1

rossicus Tshernova 1928 [Ecdyonurus]  

ruber McDunnough 1926 [Ecdyonurus]  

rubromaculata Clemens 1913 [Heptagenia]  

rubroventris Traver (in Needham & Traver & Hsu) 1935 [Heptagenia] — in Ecdyonurus/fg1

rusticalis McDunnough 1931 [Heptagenia] — in Ecdyonurus/fg1

salvini Kimmins 1934 [Heptagenia] — in Ecdyonurus/fg1

samochai Demoulin 1973 [Sigmoneuria]

scitulum Traver (in Needham & Traver & Hsu) 1935 [Stenonema]  

semicolorata Curtis 1834 [Baetis] — typus nominis Rhithrogena — in Rhithrogena/fg3

semitincta Pictet 1843 [Baetis] — in Rhithrogena/fg3

shanwangensis Zhang 1989 [Heptagenia]

 simplex Walsh 1863 [Heptagenia]

simplicioides McDunnough 1924 [Heptagenia] — typus nominis Akkarion — in Ecdyonurus/fg1

sinclairi Lewis 1979 [Stenonema]

sinex Demoulin 1968 [Heptagenia]

smithae Traver 1937 [Stenonema]

solangensis Dubey 1971 [Heptagenia]

soldatovi Tshernova 1952 [Heptagenia]  

solitaria McDunnough 1924 [Heptagenia]

spinosa Traver 1933 [Heptagenia] — in Ecdyonurus/fg1

straminea Curtis 1834 [Baetis]  

 subaequalis Banks 1914 [Heptagenia] — in Rhithrogena/fg2-Cinygmula/g1

sulphurea Müller 1776 [Ephemera]

suzukiella Matsumura 1931 [Kageronia]  

sylvicola Pictet 1865 [Baetis] — in Epeorus/fg3

tadzhikorum Tshernova 1931 [Heptagenia]  

terminata Walsh 1862 [Palingenia]

tena [Heptagenia] nomen nudum

tessellata Walker 1853 [Baetis]  

thetis Traver (in Needham & Traver & Hsu) 1935 [Heptagenia] — in Ecdyonurus/fg1

tobiironis Takahashi 1929 [Ecdyurus] — in Ecdyonurus/fg1

torrida Walker 1853 [Baetis]

townesi Traver (in Needham & Traver & Hsu) 1935 [Heptagenia]

traverae Braasch 1986 [Heptagenia]

 trimaculata Ikonomov 1963 [Heptagenia] — in Ecdyonurus/fg1

tripunctata Banks 1910 [Heptagenia]  

umbratica McDunnough 1931 [Heptagenia] — in Ecdyonurus/fg1

undulatus Banks 1924 [Epeorus] — in Rhithrogena/fg3

varium Traver 1933 [Stenonema]  

venosa Fabricius 1775 [Ephemera] — typus nominis Ecdyonurus — in Ecdyonurus/fg1

 verticis Say 1839 [Baetis] — typus nominis Anthopotamus — in Potamanthus/fg1

vicaria Walker 1853 [Baetis]

viridis Matsumura 1931 [Heptagenia] — in Ecdyonurus/fg1

vitrea Walker 1853 [Palingenia] — in Epeorus/fg3

volitans Eaton 1870 [Heptagenia]  

wabasha Daggy 1945 [Stenonema]

walshi McDunnough 1926 [Heptagenia] — in Ecdyonurus/fg1

werestschagini Tshernova 1952 [Heptagenia] — in Ecdyonurus/fg1

whitingi Webb & McCafferty (in Webb & Sun & McCafferty & Ferris) 2007 [Heptagenia]

wudigeum McCafferty & Lenat 2010 [Maccaffertium]

yoshidae Takahashi 1924 [Ecdyonurus] — in Ecdyonurus/fg1

 zebrata Hagen 1864 [Baetis] — in Ecdyonurus/fg1


Heptageniidae genn.spp.

sources:

subjects:

actual arbitrary names:

synonymous arbitrary names:

valid names:

Traver 1933

 

nameless genus

 

Ironodes

Demoulin 1955

wing

Ecdyonuridae gen.sp.

 

 

Demoulin 1964

Afghanistan

 

 

 

Wais & Campos 1984:IV.Conf.Eph.

S.America

 

 

 

Lewis & Wehr 1993'

Heptageniidae gen.sp.

 

 

Nakamura & Kameyama 
& Katayama 2001

Japan

Heptageniidae gen.sp.

 

 

Zhou & Jacobus & DeWalt & 
Adamowicz & Hebert 2010

DNA barcode

Heptageniidae sp.

 

 

Wichard 2005

Baltic amber

Heptageniidae

 

 

Bell & Fanti & Acorn et al. 2013

cf. Heptageniidae

 

 

Vuataz & Sartori & Gattoliat & Monaghan 2013

DNA

Heptageniidae sp.1

 

ornata [Rhithrogeniella]

 

 

 

 

 

 

 

 

 

 


Heptagenia spp.

sources:

subjects:

actual arbitrary names:

synonymous arbitrary names:

valid names:

Banks (in Baker 1905)

California

Heptagenia sp.n.

 

 

Needham 1905*

Heptagenia sp.1

 

 

-"-

Heptagenia sp.2

 

 

-"-

Heptagenia sp.3

 

interpunctata [Baetis

Clemens 1913*

Heptagenia sp.

 

 

Clemens 1915

Heptagenia sp.

 

 

Traver 1933:105-125*

?

Heptagenia sp.?

 

julia [Heptagenia]

Traver 1939

Heptagenia sp.A

 

traverae [Heptagenia

-"-

Heptagenia sp.B

 

 

Zhadin 1940

Kolskiy pen.

Heptagenia sp.

 

 

Imanishi 1940

Heptagenia na

 

arsenjevi [Heptagenia]

flava [Heptagenia

-"-

Heptagenia nb

 

tena [Heptagenia] nomen nudum

kyotoensis [Heptagenia

Tshernova 1947

N.Russia

Heptagenia sp.

 

 

Mayo 1952:179-186*

Heptagenia sp.1

 

 

Gose 1962

Japan

Heptagenia HB

= Heptagenia na: Imanishi 1940

?flava [Heptagenia]

-"-

Japan

Heptagenia nb

= Heptagenia nb: Imanishi 1940

kyotoensis [Heptagenia

Levanidova 1964

 

Heptagenia sp.

 

tobiironis [Ecdyurus

?dracon [Ecdyonurus]

Bajkova 1965

 

Heptagenia sp.

 

tobiironis [Ecdyurus

?dracon [Ecdyonurus]

Kasymov 1972

Caucasus

Heptagenia sp.

 

 

Puthz 1973

Israil

Heptagenia sp.1 (gr. lateralis)

 

 

-"-

Israil

Heptagenia sp.2 (gr. lateralis)

 

 

Edmunds & Jensen 
& Berner 1976

Centr.America

Heptagenia sp.?

 

 

Bajkova 1978

Mongolia

Heptagenia sp.

 

 

Lewis 1978*

OLIGOCENE

Heptagenia sp.

 

 

Gose 1979*

Japan

Heptagenia sp.na

= HB

 

Landa & Soldan 1983

Mongolia

Heptagenia sp.

 

 

Gose 1985

Japan

Heptagenia HB

= Heptagenia na: Imanishi 1940

?flava [Heptagenia]

Kluge 1987

Pamirs

Heptagenia sp.

 

quadripunctata [Heptagenia

Zhang 1989

MIOCENE, China

Heptagenia sp.

 

 

Soldan 1991

Vietnam

Heptagenia sp.

 

 

Nakamura & Kameyama 
& Katayama 2001

Japan

Heptagenia sp.

 

 

 

Heptagenia sp.

 

 

 

Heptagenia sp.

 

 

 

Heptagenia sp.