CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Plesiomorphon Torleya/g2
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Furcatergaliae
Ephemerella/fg1Tricorythus/fg1Ephemerella/fg2Timpanoga/fg1Ephemerella/fg3
Hyrtanella/f1-Torleya/g1
- pm.Torleya/g2)
Nomen hierarchicum:
Torleya/g2
[g:1917] (sine Teloganopsis, Crinitella, Uracanthella, Amurella;
incl. Serratella)
In circumscription fits:
— plesiomorphon Torleya/g2: Kluge 2004: 312
Nominal taxa included:
Serratella/g
[g:1959]
Vittapallia/g [g:2008]
Quatica/g [g:2008]
References.
Allen & Edmunds 1963b (Serratella): 
'
'; – Kluge 1997: *;
– Kluge 2004: 
* *
* *
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Plesiomorphies of Torleya/g2. Unlike Teloganopsis and Hyrtanella/f2=Crinitella/g1, hind wing without prominent costal projection and with Sc terminating near apex (Kluge 2004: Fig.90:B) (as in all other Ephemerella/fg2). Unlike Uracanthella, maxilla retains biting structure with well-developed canines (Ikonomov 1961: Fig.3:13). Unlike Amurella, larval abdominal terga either with paired projections [see Ephemerella/fg1 (19)], or without projections, but never have unpaired projections. Variable characters of Torleya/g2. Tergalii III [anteriormost — see Ephemerella/fg3 (1)] can have normal size, or more or less strongly enlarged and form gill opercula covering the rest pairs of tergalii; the same variability in Hyrtanella/f2=Crinitella/g1. |
Size. Fore wing length 5–10 mm.
Distribution. Holarctic and Oriental Region.
Nominal species in Torleya/g2:
acutiformis Zhou (in Ding & Jacobus & Zhou) 2022 [Serratella]
aeneus Pictet 1843 [Potamanthus]
— syn.subj. ignita [Ephemera]
altana Allen 1968 [Ephemerella]
— syn.subj. micheneri [Ephemerella]
andaluciaca
Kazanci 1990 [Drunella]
— syn.subj.
ikonomovi [Ephemerella]
angusta Traver 1934 [Ephemerella]
— syn.subj. tibialis
[Ephemerella]
angustata Berner 1950 [Ephemerella]
— syn.subj. tibialis
[Ephemerella]
antuensis Su & You 1989 [Ephemerella (Ephemerella)]
— syn.subj. ignita [Ephemera]
arenosa Tong & Dudgeon 2000 [Torleya]
— syn.subj.
nepalica [Ephemerella (Torleya)]
belgica Lestage 1917 [Torleya]
— typus nominis Torleya
— syn.subj. major [Ephemerella]
brevicauda Jacobus & Zhou & McCafferty 2009 [Serratella]
carolina Berner & Allen 1961 [Ephemerella (Serratella)]
— syn.subj. serrata [Ephemerella]
chantauense
[Ephemerella]
(in litt.)
,
,
-
-
/
cornutus Gose 1980 [Ephemerella (Ephemerella)]
nom.praeocc.
— syn.obj. tsuno [Serratella]
dibruensis
Selvakumar & Martynov & Jacobus (in Martynov & Selvakumar
& Subramanian & Sivaramakrishnan & Vasanth & Sinha & Jacobus
2021) [Torleya]
diluta Stephens 1835 [Ephemera]
— syn.subj. ignita [Ephemera]
elissa Jacobus & Zhou & McCafferty 2009 [Serratella]
erythrophthalma Schrank 1798 [Ephemera]
— syn.subj. ignita [Ephemera]
frisoni McDunnough 1927 [Ephemerella]
gibbus Pictet 1843 [Potamanthus]
— syn.subj. ignita [Ephemera]
glareosa Kang & Yang 1995 [Torleya]
— syn.subj. nepalica [Ephemerella (Torleya)]
grandipennis Zhou & Su & Hong 2000 [Torleya]
ignita Poda 1761 [Ephemera]
--/
ikonomovi Puthz 1971 [Ephemerella]
— typus nominis Quatica
imanishii Gose 1980 [Ephemerella (Ephemerella)]
nom. praeocc. —
syn.obj. occiprens [Serratella]
ishiwatai Gose 1980 [Ephemerella (Ephemerella)]
japonica Gose 1980 [Ephemerella (Torleya)]
karasuensis Kustareva 1976 [Drunella]
— syn.subj. ignita [Ephemera]
karia Kazanci 1990 [Drunella]
kogistana [Ephemerella]
(in litt.)
--/
lactata Bengtsson 1909 [Ephemerella]
— syn.subj. ignita [Ephemera]
lacuna Jacobus & McCafferty & Sites 2007 [Crinitella]
leonidi Martynov & Palatov (in Martynov & Palatov &
Godunko) 2023 [Serratella]
--/
levis Day 1954 [Ephemerella]
lutosa Kang & Yang 1995 [Torleya]
maculocaudata Ikonomov 1961 [Ephemerella]
--/
major Klapálek
1905 [Ephemerella]
,,
micheneri Traver 1934 [Ephemerella]
-
mikhaili
Tiunova 1995 [Torleya]
,
naga Jacobus & McCafferty (in Jacobus & Zhou & McCafferty) 2004
[Torleya]
nazarita Alba-Tercedor & Derka
2003 [Torleya]
nepalica Allen & Edmunds 1963 [Ephemerella (Torleya)]
nigrosetosa Bengtsson 1917 [Ephemerella torrentium]
— syn.subj. ignita [Ephemera]
nuda Tshernova 1949 [Ephemerella]
--/
obscura
Stephens 1835 [Baetis]
— syn.subj. ignita [Ephemera]
obscura Bengtsson 1917 [Ephemerella torrentium]
— syn.subj. ignita [Ephemera]
occiprens
Jacobus & McCafferty 2008 [Serratella]
padunica
Kazlauskas 1963 [Ephemerella (Torleya)]
,-
palatovi
Martynov & Selvakumar & Jacobus (in Martynov & Selvakumar
& Subramanian & Sivaramakrishnan & Vasanth & Sinha & Jacobus
2021) [Serratella]
paradinasi Gonzalez-del-Tanago
& Garcia-de-Jalon 1981 [Drunella]
-
rosea Stephens 1835 [Ephemera]
— syn.subj. ignita [Ephemera]
rufescens
Stephens 1835 [Ephemera]
— syn.subj. ignita [Ephemera]
rufescens
Bengtsson 1917 [Ephemerella torrentium]
— syn.subj. ignita [Ephemera]
sequoia Allen & Collins 1968 [Ephemerella]
— syn.subj. tibialis
[Ephemerella]
serrata Morgan 1911 [Ephemerella]
— typus nominis Serratella
serratoides
McDunnough 1931 [Ephemerella]
,-
setigera Bajkova 1965 [Ephemerella]
--/
sibirica Tshernova 1952 [Ephemerella]
— syn.subj. ignita [Ephemera]
simbalbarensis
Selvakumar & Subramanian & Martynov & Jacobus (in Martynov & Selvakumar
& Subramanian & Sivaramakrishnan & Vasanth & Sinha & Jacobus
2021) [Torleya]
sordida McDunnough 1925 [Ephemerella]
— syn.subj. serrata [Ephemerella]
spiculosa Berner & Allen 1961 [Ephemerella]
— syn.subj. serrata [Ephemerella]
spinosa Ikonomov 1961 [Ephemerella]
nom.praeocc.
— syn.obj.
ikonomovi [Ephemerella]
subsolana
Allen 1973 [Ephemerella (Serratella)]
thymalli
Tshernova 1952 [Ephemerella]
--/
tibialis
McDunnough 1924 [Ephemerella]
— typus nominis Vittapallia
torrentium Bengtsson 1917 [Ephemerella]
— syn.subj. ignita [Ephemera]
tsuno
Jacobus & McCafferty 2008 [Serratella]
tumiforceps Zhou & Su 1997 [Serratella]
— syn.subj. nepalica [Ephemerella (Torleya)]
uenoi Allen & Edmunds 1963 [Ephemerella (Drunella)]
undatella Allen 1971 [Ephemerella (Acerella)]
— syn.obj. uenoi [Ephemerella (Drunella)]
variegata Bengtsson 1917 [Ephemerella torrentium]
— syn.subj. ignita [Ephemera]
verrucosa Kluge 1980 [Ephemerella]
--/
zapekinae
Bajkova 1967 [Ephemerella]
--/
See also:
Torleya/g1 INCERTAE SEDIS
Ephemerella/fg3 INCERTAE SEDIS
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