CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Asthenopus/fg1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Furcatergaliae
Fimbriatotergaliae
Fossoriae
Cryptoprosternata
Polymitarcys/f1=Ephoron/g2
Campsurus/fg1
- Asthenopus)
Nomen hierarchicum:
Asthenopus/fg1
[f:1954; g:1871] (incl. Povilla, Asthenopodes, Languidipes)
In circumscription fits:
— subfam. Asthenopodinae Edmunds & Traver 1954a:239
— Asthenopus/fg1: Kluge 2004: 266
Nominal taxa included:
Povilla/g
[g:1912]
Asthenopodes/g
[g:1924]
Languidipes/g
[g:1984]
References. Ulmer 1916 (Povilla): 
'; – Needham & Murphy 1924:
'; – Ulmer 1939 (Povilla):
(L)'; – 1942: ; – Spieth 1943:
; – Traver 1950: ; – Demoulin
1956c (Povilla): 
'; – 1966a:
; – Edmunds & Allen &
Peters 1963: *; – Roback 1966:
'; – Hubbard 1984 (Povilla):
' '; – Dominguez 1988:
; – Hubbard & Dominguez 1988:
; –
Kluge 2004: 
* *
* *
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Autapomorphies of Asthenopus/fg1. (1) Mandibular tusks [see Campsurus/fg1 (1)] are specialized as biting: not long, very thick and stout, with serrate inner margin; these tusks resemble mandibular incisors of other biting insects (Kluge 2004: Fig.79:C–D). (2) Larval claw of fore leg [see Campsurus/fg1 (6)] has a row of denticles on inner margin (while claws of middle and hind legs have no denticles). Unique apomorphy: in all other Fossoriae denticles on all claws are absent; in that Ephemeroptera where denticles are present, they are developed on all legs. Plesiomorphies of Asthenopus/fg1. On fore wing CuA is not so strongly curved as in Campsurus/fg2, thus both intercalaries of cubital field [see Campsurus/fg1 (5)] go nearly parallel to basitornal margin and terminate near or anteriad of tornus (Kluge 2004: Fig. 79:A). Unlike Campsurus/fg2, genitals retain small median styliger, which is articulated to posterior margin of sternite IX and bears immobile gonostyli pedestals [narrow basally – see Polymitarcys/f1=Ephoron/g2 (7)]; penial arms retain lateral articulations with posterior-lateral angles of tergite IX; gonostyli pedestals retain muscles, which move gonostyli [non-segmented – see Campsurus/fg1 (10)]. In spite of short-living specialization of adult [see Polymitarcys/f1=Ephoron/g2 (3)], moult from subimago to imago takes place not only in male, but in female as well. Unlike Campsurus/fg2, egg has a usual ellipsoid shape; long threads being coiled, can either form two polar caps, or cover the whole egg surface (Koss & Edmunds 1974: Fig.211–214). Vestige of tergalius I [see Cryptoprosternata (6)] is bilamellate. |
Size. Fore wing length 8–20 mm.
Distribution. Afrotropical, Oriental and Neotropical Regions.
Nominal species in Asthenopus/fg1:
adusta Navás
1912 [Povilla]
— typus nominis Povilla
,
,
typus
amazonicus Hagen 1888 [Campsurus]
— syn.subj. curta
[Palingenia]
andamanensis Hubbard 1984 [Povilla
(Povilla)]
angelae
De-Souza & Molineri 2012 [Asthenopus] 
cambodjensis Ulmer 1920 [Povilla]
chumuco Molineri &
Salles & Peters 2015 [Asthenopodes]
corporaali
Lestage 1922
[Asthenopus]
crenulatus
Molineri & Cruz & Emmerich 2011 [Asthenopus]
— typus nominis Hubbardipes
curta
Hagen 1861 [Palingenia]
— typus nominis Asthenopus
exuv.;,
-,
//
gilliesi Domínguez 1988 [Asthenopus]
— typus nominis Priasthenopus
guarani
Molineri &
Salles & Peters 2015 [Asthenopus]
heardi Hubbard 1984 [Povilla
(Povilla)]
,,
,
hubbardi
Molineri &
Salles & Peters 2015 [Asthenopus]
janae
Hankel & Molineri 2024 [Languidipes]
junki Hubbard 1984 [Povilla
(Povilla)]
— syn.subj. heardi [Povilla]
lithophagus
Bolotov & Kondakov & Potapov & Palatov & Spitsyn (in Bolotov
& Kondakov & Potapov & Palatov & Lunn & Bovykina &
Chapurina & Kolosova & Spitsyna & Spitsyn & Lyubas & Gofarov
& Vikhrev & Yapaskurt & Bychkov & Pokrovsky) 2022 [Languidipes]
magnus
Molineri &
Salles & Peters 2015 [Asthenopus]
paraguarius
Navás 1920 [Campsurus]
picteti Hubbard 1975 [Asthenopodes]
taprobanes Hubbard 1984 [Povilla
(Languidipes)]
traverae Molineri &
Salles & Peters 2015 [Asthenopodes]
ulmeri
Hubbard 1984 [Povilla (Povilla)]
— syn.subj. heardi [Povilla]
See also:
Campsurus/fg1
INCERTAE SEDIS
