CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Ameletus/fg2
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
pm.Tridentiseta
Ameletus/fg1
- Ameletus/fg2)
Nomen hierarchicum:
Ameletus/fg2
[f:1991; g:1885] (sine Metreletus)
In circumscription fits:
— gen. Ameletus Eaton 1885: 210
— Ameletus/fg2: Kluge 2004: 82
Nominal taxa included:
Chimura/g
[g:1915]
Paleoameletus/g
[g:1940]
References. Eaton 1883-1885: 
*
*; – Needham & Traver & Hsu
1935: * *; –
Edmunds & Jensen & Berner 1976:
* *; –
Zloty
1996: * *; –
Kluge 2004:
* 
*
*; – Kluge 2007:
* * *.
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Autapomorphies of Ameletus/fg2. (1) Glossae [see Ameletus/fg1 (1)] are truncate, i.e. with widened straight apical margin; all apical setae form a regular apical-ventral row and are flat, widened toward apex and rounded apically (Kluge 2004: Fig.20:A). Unique apomorphy. Characters of Ameletus/fg2 of unclear phylogenetic status. (2) Larval protopenis has simple shape: each lobe is triangular or conic, with rounded apex and without incision between lateral lobe and ventral plate [which are present on imaginal penis – see Ameletus/fg1 (6)] (Kluge 2007: Fig.10, 21); apex of this larval penis lobe corresponds to the apex of imaginal lateral lobe. This shape of larval penis is retained not only in species whose imaginal lateral lobes are long, but also in oregonensis [Ameletus] whose imaginal lateral lobes are shorter than median plates and strongly divergent. Plesiomorphies of Ameletus/fg2 (unlike Metreletus). Cubital field of fore wing without intercalaries, several (4–8) veins go from CuA to basitornal margin [see Anteritorna (1)]. Larval claws have no denticles. |
Size. Fore wing length 7–17 mm.
Distribution. Holarctic and Oriental Region, dominate in Nearctic and Eastern Palaearctic; in Western Palaearctic occurs a single Holarctic species – inopinatus [Ameletus].
Nominal species in Ameletus/fg2:
aequivocus McDunnough 1934
[Ameletus]
— syn.subj. sparsatus
[Ameletus]
aetherea Navás 1915 [Chimura]
— typus nominis
Chimura
alexandrae Brodsky
1930 [Ameletus]
--/
allengaensis Tiunova (in Tiunova & Semenchenko &
Veyaev) 2017 [Ameletus] 
alpinus Bengtsson 1913 [Ameletus]
— syn.subj. inopinatus [Ameletus]
altaicus
Kluge 2007 [Ameletus]
--/
alticolus McDunnough 1934 [Ameletus]
— syn.subj. celer [Ameletus]
amador Mayo 1939 [Ameletus]
andersoni Zloty 1996 [Ameletus]
arlecchino
Kluge 2007 [Ameletus montanus] --/
asiacentralis Soldán 1978 [Ameletus]
— syn.subj. alexandrae [Ameletus]
atratus Kang & Yang 1994 [Ameletus]
bellulus
Zloty 1996 [Ameletus]
-
browni
McDunnough 1933 [Ameletus]
camtschaticus
Ulmer 1927
[Ameletus]
--/
cedrensis
Sinitshenkova 1977 [Ameletus]
--/
celer McDunnough 1934 [Ameletus]
-
celeroides McDunnough 1934
[Ameletus]
— syn.subj. celer [Ameletus]
connectina
McDunnough 1939
[Ameletus]
— syn.subj. velox [Ameletus]
connectus McDunnough 1936 [Ameletus]
— syn.subj. velox [Ameletus]
cooki
McDunnough 1929 [Ameletus]
costalis Matsumura 1931 [Chimura]
,-
,/
cristatus Bajkova 1976 [Ameletus]
crocerus
Imanishi 1931 [Ameletus]
cryptostimulus Carle 1978 [Ameletus]
daliensis
Tong (in Li & Luo & Jiang & Wang & Tong) 2001 [Ameletus]
dissitus
Eaton 1885 [Ameletus]
doddsianus Zloty 1996 [Ameletus]
dodecus
Dubey 1971 [Ichthybotus]
edmundsi
Zloty 1996 [Ameletus]
eugenii Sinitshenkova &
Varykhanova 1989 [Ameletus]
— syn.subj. inopinatus [Ameletus]
exquisitus Eaton 1885 [Ameletus]
facilis Day 1952 [Ameletus]
— syn.subj. vancouvernensis [Ameletus]
falsus McDunnough
1938 [Ameletus]
formosus Kang & Yang 1994 [Ameletus]
imbellis Day 1952 [Ameletus]
inopinatus Eaton 1887 [Ameletus]
--/
janetae Kondratieff & Meyer 2010 [Ameletus]
khasanensis Tiunova 2013 [Ameletus]
kyotensis Imanishi 1932 [Ameletus]
labiatus
Sinitshenkova 1981
[Ameletus]
--
lineatus Traver
1932 [Ameletus]
longulus Sinitshenkova 1981 [Ameletus]
,-,/
ludens Needham 1905 [Ameletus]
majusculus Zloty 1996 [Ameletus]
minimus Zloty
& Harper 1999 [Ameletus]
monta
Mayo 1952 [Ameletus]
— syn.subj. similor [Ameletus]
montanus Imanishi 1930 [Ameletus]
--/
montivagus Kang & Yang
1994 [Ameletus]
oregonensis
McDunnough 1933 [Ameletus]
-
parvus Kluge 1979 [Ameletus]
--/
pilatus Sinitshenkova 1981 [Ameletus]
— syn.subj. camtschaticus
[Ameletus]
primitivus Traver
1939 [Ameletus]
— typus nominis Paleoameletus
pritchardi Zloty 1996 [Ameletus]
procerus Bajkova 1976 [Ameletus]
quadratus
Zloty & Harper 1999 [Ameletus]
querulus McDunnough 1938 [Ameletus]
— syn.subj. shepherdi [Ameletus]
rossicus
Kluge 2007 [Ameletus montanus]
--/
shepherdi Traver 1934 [Ameletus]
similor McDunnough 1928 [Ameletus]
/,
sirotskii Tiunova (in Tiunova & Semenchenko & Veyaev)
2017 [Ameletus]
sparsatus
McDunnough 1931 [Ameletus]
subalpinus Imanishi 1931 [Ameletus]
subnotatus Eaton 1885 [Ameletus]
— typus nominis Ameletus
suffusus McDunnough
1936 [Ameletus]
tarteri Burrows 1987 [Ameletus]
tertius McDunnough
1938 [Ameletus]
tolae Zloty 1996 [Ameletus]
tuberculatus McDunnough 1939 [Ameletus]
— syn.subj. celer [Ameletus]
validus McDunnough 1923 [Ameletus]
vancouvernensis McDunnough 1933 [Ameletus]
velox Dodds 1923 [Ameletus]
-
vernalis McDunnough 1924 [Ameletus]
walleyi Harper
1970 [Ameletus]
Examined also:
sp. (China) /
