CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Afronurus/fg1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Branchitergaliae
Heptagennota
Pentamerotarsata
Radulapalpata
Heptagenia/f5=g4
Ecdyonurus/fg1
Atopopus/fg1
- Afronurus/g1)
Nomen hierarchicum:
Afronurus/fg1
[f:2007; g:1924] (incl. Cinygmina)
In circumscription fits:
— Afronurus/g1: Kluge 2004: 184
Nominal taxon included:
Cinygmina/g
[g:1937]
References. Koss & Edmunds 1974:
*; –
Kluge 2004: * 
*
* 
*
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Autapomorphies of Afronurus/fg1. (1) Median titillators of penis [see Pentamerotarsata (5)] are reduced: they are either widely separated and diminished (in assamensis [C.], yoshidae [E.], levis [E.]), or completely lost. This apomorphy is non-unique among Pentamerotarsata, but does not occur in other Heptagenia/f5=g4 (which have median titillators well-developed and contiguous medially). (2) Egg [bearing small anchors – see Branchitergaliae (4)] on equator bears several additional anchors of another structure: the additional anchor is many times larger than a usual anchor, and in coiled condition represents a flat spiral (Koss & Edmunds 1974: Fig.94–95; Kopelke 1980: Fig.26–29; Flowers & Pescador 1984: Fig.11–12; Kang & Yang 1994: Fig.17–18). Unique apomorphy. Among Afronurus/g1 such additional anchors are present not in all species: among species examined, they are present in levis [E.], spp. cf. assamensis [C.] and peringueyi [E.]; are reported for chihpenensis [A.], hyalinus [E.], philippinensis [A.] and viridis [E.]; but these additional anchors are absent in abracadabrus [E.] (at the same time abracadabrus [E.] is attributed to Afronurus/fg1 on the base of great similarity with levis [E.] in larval structure). |
Size. Fore wing length 7–13 mm.
Distribution. Africa and Asia. Possibly in Miocene was distributed in Europe as well, i.e. had Arctogean distribution (see Pseudokageronia in Ecdyonurus/fg1 INCERTAE SEDIS)
Nominal species in Afronurus/fg1:
abracadabrus Kluge 1983 [Ecdyonurus]
--/
aethereus Navás
1936 [Ecdyonurus]
alces
Braasch & Jacobus 2011
[Afronurus]
assamensis
Kimmins 1937 [Cinygmina]
— typus nominis Cinygmina
//,/
barnardi Schoonbee 1968 [Afronurus]
--/
cervina Braasch & Soldán
1984 [Cinygmina]
--/
chihpenensis Kang & Yang 1994 [Afronurus]
collarti
Navás 1930 [Adenophlebia]
dama Braasch & Soldán 1987 [Cinygmina]
--/
drepanophyllus Zhang & Lei & Li & Zhou 2021 [Afronurus]
—
syn.subj. dama [Cinygmina]
elgonensis Puthz 1971 [Afronurus]
floreus
Kang & Yang 1994 [Afronurus]
freitagi Braasch
2011
[Afronurus]
furcata Zhou & Zheng 2003 [Cinygmina]
gilliesi
Corbet 1962 [Afronurus]
gilliesiana
Braasch 1990 [Cinygmina]
hainanensis She & Gui & You 1995
[Cinygmina]
— syn.subj.
rubromaculata [Cinygmina]
harrisoni Barnard 1932 [Afronurus]
hunanensis
Zhang & Cai 1991
[Cinygmina]
hyalinus Ulmer 1912 [Ecdyurus]
japonicus Ueno 1928 [Ecdyonurus]
— syn.subj. yoshidae [Ecdyonurus]
keralensis
Braasch & Soldán 1987 [Cinygmina]
--/
kiangsuensis
Puthz 1971 [Ecdyonurus]
kumbakkaraiensis Venkataraman & Sivaramakrishnan 1989 [Cinygmina]
— syn.subj. keralensis [Cinygmina]
landai Braasch &
Soldán 1984 [Cinygmina]
lantuyanensis
Braasch 2011 [Afronurus]
levis Navás 1912 [Epeorus]
--/
leytenensis Braasch 2011
[Afronurus]
malaysianus Braasch 2005
[Afronurus]
meenmutti Muthukatturaja & Balasubramanian 2021 [Afronurus]
— syn.subj. keralensis [Cinygmina]
mindanaoensis Braasch 2011
[Afronurus]
mindoroensis Braasch 2005
[Afronurus]
muehlenbergi Puthz 1971 [Afronurus]
namnaoensis Braasch & Boonsoong 2010 [Afronurus]
— syn.subj. dama [Cinygmina]
nanhuensis
Kang & Yang1994 [Afronurus]
negi
Corbet 1960 [Afronurus]
obliquistriata You & Tian
& Hong & Hsu 1981 [Cinygmina]
— syn.subj. pallescens
[Ecdyonurus]
oliffi Schoonbee 1968 [Afronurus]
otus
Braasch & Jacobus 2011
[Afronurus]
panayensis Braasch
2011
[Afronurus]
pallescens
Navás 1936 [Ecdyonurus]
peringueyi Esben-Petersen
1913 [Ecdyonurus]
— typus nominis Afronurus
/
philippinensis Flowers
& Pescador 1984 [Afronurus]
pulcher Ulmer
1930 [Afronurus]
— syn.subj. collarti
[Adenophlebia]
rainulfiana
Braasch 1990 [Cinygmina]
rangifera Braasch & Soldán
1987 [Cinygmina]
rubromaculata You & Wu & Gui & Hsu
1981 [Cinygmina]
-
-
/
sarawakensis Braasch
2011
[Afronurus]
scotti Schoonbee 1968 [Afronurus]
subflavus
Kopelke 1980 [Afronurus]
ugandanus Kimmins 1956 [Afronurus]
viridis
Matsumura 1931 [Heptagenia]
webbi Braasch
2011
[Afronurus]
yixingensis
Wu & You 1986 [Cinygmina]
yoshidae Takahashi 1924 [Ecdyonurus]
,
zachvatkini Tshernova 1952 [Cinygmula]
— syn.subj. levis [Epeorus]
Examined also:
sp.
(Thailand) --/
spp. (Africa)
See also:
Ecdyonurus/fg1 INCERTAE SEDIS
Radulapalpata INCERTAE SEDIS
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