EPHEMEROPTERA OF THE WORLD

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hierarchical names based on Baetis

Baetoidea

Baetidae

Baetinae

Baetini

genus Baetis

subgenus Baetis

sources:

subjects:

actual arbitrary names:

synonymous arbitrary names:

valid names:

Eaton 1868

Baetis sect.A

Centroptilum

-"-

Baetis sect.B

-"-

Baetis sect.C

Eaton 1883-1888

Peru: Puno

! Cloeodes sp.

Ulmer 1904

fuegiensis [Ameletus]

anceps [Siphlurus]

Neizvestnova-Zhadina 1931

r.Oka

shadini [Pseudocentroptilum]

macronyx [Cloeon (Procloeon)]

Traver 1944

-"-

-"-

-"-

Kimmins (1955)1956*

Baetidae A, nr.Centroptilum

berneri [Afrobaetodes]

Roback 1966

Dactylobaetis

Camelobaetidius

Traver & Edmunds 1968

Peru

Baetidae gen.unknown

Allen & Murvosh 1983*

USA

peninsulus [Cloeodes]  

Müller-Liebenau 1984*

Malaysia

continentale [Indocloeon (Hindocloeon)] 

-"-

Malaysia

Bungona (Chopralla) sp.

Crassolus sp.

Wolda & Flowers 1985

Edmunds & Polhemus 1990

N'Guinea

gen.A

-"-

Celebes

gen.E

Echinobaetis

-"-

Celebes

gen.G

-"-

Celebes

Baetis-ally

Flowers 1990*

Fuji

-"-

Fuji

Flowers 1991*

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

-"-

Panama

Suter 1992

gen.A sp.1

gagadjuensis [Platybaetis]

Pescador & Masteller 
& Buzby 1993

Puerto Rico

Dean & Suter 1996*

gen.1

= Baetis (partim)

-"-

gen.2

= Baetis (partim)

-"-

gen.3

= Pseudocloeon (partim)

-"-

gen.4

= Baetis (partim)

-"-

gen.5

= Pseudocloeon (partim)

Suter 1997

soror [Baetis] 

-"-

-"-

-"-

-"-

confluens [Baetis]

...

-"-

-"-

sobrinus [Offadens] 

-"-

frater [Baetis]

-"-

-"-

-"-

hickmani [Edmundsiops] 

-"-

-"-

-"-

baddamsae [Baetis] 

-"-

-"-

inconspicuum [Pseudocloeon] 

-"-

plectile [Pseudocloeon] 

-"-

hypodeum [Pseudocloeon] 

McCafferty 1997

Lebanon amber

Monaghan & Gattolliat & 
Sartori & Elouard & James & 
Derleth & Glaizot & de Moor 
& Vogler 2005

rDNA

Gattolliat & Sartori 2006

Ghana

Gattolliat & Monaghan & Sartori 
& Elouard & Barber-James 
& Derleth & Glaizot & de Moor 
& Vogler 2008:XI.Conf.Eph*

rDNA

Gen.nov.

Zhou & Jacobus & DeWalt & 
Adamowicz & Hebert 2010

DNA barcode

Baetidae sp.

Falcao & Salles & Hamada 2011

Brazil photo!

Baetidae gen.sp.nov.

tunamore [Macuxi]

Cruz & Salles & Hamada 2018

Brazil 

Baetidae gen.sp.A

tunamore [Macuxi]

Baetidae gen.sp.

= Hagenulodes braueri: Gerlach 2001

sources:

subjects:

actual arbitrary names:

synonymous arbitrary names:

valid names:

Pictet 1846

Baetis sp.

Walker 1853

Baetis?

ignava [Baetis]

Eaton 1883-1888

Hindostan

Baetis —.

= Cloeon debilis: 
Walker 1860

= Baetis debilis: Eaton 1871 
(non Baetis debilis Walker 1853)

debilis [Cloeon]

Bengtsson 1904

Baetis sp.n.

-"-

Baetis sp.n.

-"-

Baetis sp.n.?

Esben-Petersen 1908

 Norvegia

Baetis sp.

Esben-Petersen 1912

Argentina

Baetis sp.

Esben-Petersen 1916

  Polar Urals

Baetis sp.1

-"-

  Polar Urals

Baetis sp.2

Lestage 1917

Baetis sp.

Lestage 1917:121-140

Baetis sp.

-"-

Baetis sp.

-"-

?[Baetis sp.]

-"-

Baetis sp.

Needham & Murphy 1924*

Peru

Baetis sp.? No.1

-"-

Peru

Baetis sp.? No.2

Seeman 1927

  California

Baetis sp.1

-"-

  California

Baetis sp.2

-"-

  California

Baetis sp.3

Ulmer 1927

 Kamchatka

Baetis sp.

Neizvestnova-Zhadina 1930

 r.Oka

Baetis sp.

Traver 1932*

Baetis sp.1

incertans [Baetis] 

tricaudatus [Baetis] 

-"-

Baetis sp.

Ueno 1935

  Japan

Baetis sp.

Traver 1938

  Porto Rico

Baetis sp.1

-"-

  Porto Rico

Baetis sp.2

Tshernova 1938

 Caucasus

Baetis sp.

Wingfield 1939

tergalii 

Baetis sp.

Imanishi 1940

Baetis n1a

fuscata [Ephemera]  

-"-

Baetis n1b

-"-

Baetis n1c

-"-

Baetis nX

sibiricum [Pseudocloeon] 

Zhadin 1940

 Caucasus

Baetis sp.1

-"-

 Caucasus

Baetis sp.2

Tshernova 1941

r.Ili  

Baetis sp.

Traver 1944*

Brazil 

Baetis sp.

Landa 1945

 Czechoslovakia

Baetis sp.

Tshernova 1949

 r.Biya

Baetis sp.1

-"-

 r.Biya

Baetis sp.2

-"-

 r.Biya

Baetis sp.3

Baetis sp.

Gillies 1949

India

Baetis sp.1

-"-

India

Baetis sp.2

-"-

India

Baetis sp.3

Verrier 1953

Baetis sp.

Ueno 1955*

Nepal

Baetis sp.1

-"-

Nepal

Baetis sp.2

uenoi [Platybaetis]

Obr 1955

Baetis sp.

alpina [Cloe]

-"-

Baetis sp.1

melanonyx [Cloe]

Demoulin 1955

  Brazilia 

Baetis sp.

Kimmins (1955)1956*

Baetis sp.

-"-

Baetis sp.

-"-

Baetis sp.A

tsitsa [Acanthiops]

erepens [Afroptilum] 

zomba [Acanthiops]

Demoulin 1956

  Tanganjica

Baetis sp.A

-"-

  Tanganjica

Baetis sp.B

Tabacaru 1956*

Rumania

Baetis sp. (cf. rhodani)

Allen & Edmunds 1956*

 Oregon

Baetis sp.n.

Gose 1958

Japan

Baetis sp.

Verrier 1958

Mt.Nimba

Baetis sp.

Harrison & Elsworth 1958

Baetis sp.A

-"-

Baetis sp.B

latus [Baetis]

Harrison 1958

Baetis sp.A

-"-

Baetis sp.B

latus [Baetis]

Oliff 1960, 1960

Baetis sp.1

glaucus [Baetis]   

-"-

Baetis sp.2

latus [Baetis]

Allanson 1961

Baetis sp.A

-"-

Baetis sp.B

latus [Baetis]

Harrison & Agnew 1962

Baetis sp.A

glaucus [Baetis]

-"-

Baetis sp.B

latus [Baetis]

Sukackiene 1962*

  r.Angara

Baetis sp.

Chutter 1963

Baetis sp.1

= Baetis sp.1: Oliff1960

= Baetis sp.A: Harrison & Elsworth 1958

-"-

Baetis sp.2

= Baetis sp.2: Oliff1960

= Baetis sp.B: Harrison & Elsworth 1958

Agnew 1963*

 S.Africa

Baetis sp.A: Demouloin 1956

Gose 1964*

  Congo

Baetis sp.CBA

imanishii [Dicentroptilum] 

-"-

  Congo

Baetis sp.CBB

gorillora [Tanzaniella] 

Demoulin 1964

Baetis sp.1

-"-

Baetis sp.2

-"-

Baetis sp.3

-"-

Baetis sp.4

-"-

Baetis sp.5

-"-

Baetis sp.6

demoulini [Glossidion] (partim)

Levanidova 1964*

Far-East.Russia

Baetis sp.N1 (gr.rhodani)

-"-

Far-East.Russia

Baetis sp.N2 (gr.chinanonis)

Demoulin 1965

  Turkey

Baetis sp.7

-"-

  Turkey

Baetis sp.8

-"-

  Turkey

Baetis sp.9

Demoulin 1965

Baetis sp.1

-"-

Baetis sp.2

-"-

Baetis sp.3

-"-

Baetis (Acentrella) sp.4

Tanzaniella sp.

-"-

Baetis (Acentrella) sp.5

-"-

Baetis (Acentrella) sp.A

Bocharova-Messner 1965

 ontog.thorax

Baetis sp.

Roback 1966

Peru

Baetis sp.1

ellenae [Baetis] typ.nom. Mayobaetis

-"-

Peru

Baetis sp.2

-"-

Peru

Baetis sp.3

Madsen 1966

  Denmark

Baetis sp.

Grandi 1966

 Italia

Baetis sp.

Levanidova 1968

Baetis sp.1

-"-

Baetis sp. gr.vernus

-"-

Baetis sp. gr.thermicus

Ulfstrand 1968

  Lapland

Baetis sp.

Demoulin 1968

  Madagascar

Baetis sp.

in gr. molawinensis

in subgr. sumigarensis

fabulosus [Labiobaetis]

Ueno 1969

Baetis sp.1

iriomotensis [Baetis] 

-"-

Baetis sp.2

uenoi [Baetis] 

-"-

Baetis sp.3

Demoulin 1969

Baetis sp.1

-"-

Baetis sp.2

in gr. molawinensis

Demoulin 1969

  Senegal

Baetis sp.

Demoulin 1970

Baetis sp.10

-"-

Baetis sp.11

Riek 1970

wing

Baetis sp.

Samocha 1972

 Israel

Baetis  L56

soldani [Cheleocloeon] 

Levanidova 1972

Kamchatka

Baetis sp.5

Schmidt 1974

Baetis sp.

Müller-Liebenau & 
Savolainen 1975

 Finland

Baetis sp.

Bergman & Hilsenhoff 1978

Baetis sp.A

Baetis sp.B

Baetis sp.C

Bajkova 1978

 Mongolia

Baetis sp.

-"-

 Mongolia

Baetis sp.1

-"-

 Mongolia

Baetis sp.2

Morihara & McCafferty 1979:139-221*

Baetis sp.A (gr.rhodani)

adonis [Baetis] 

caelestis [Baetis]

-"-

Baetis sp.B (gr.rhodani)

libos [Baetis] 

magnus [Baetis] 

-"-

Baetis sp.C (gr.fuscatus)

notos [Baetis] 

Filipovic 1979

 Serbia

Baetis sp.

Allen 1979*

Baetis sp.

Kluge 1979

 Taimyr

Baetis sp.

Berner 1980

Baetis sp.A

-"-

  Bolivia

Baetis sp.B

incus [Bernerius]

Roldan 1980

  Colombia

Baetis sp.1

-"-

  Colombia

Baetis sp.2

-"-

  Colombia

Baetis sp.3

-"-

  Colombia

Baetis sp.4

Kopelke 1980

Baetis sp.1

Souza (1978)1981*

  Brazil

Baetis sp.B

-"-

  Brazil

Baetis sp.C

Demoulin 1981'

Baetis sp.

! Labiobaetis sp.

Dudgeon 1982:1-32

Hong Kong

Baetis T₁

-"-

Hong Kong

Baetis T₂

-"-

Hong Kong

Baetis T₃

Dudgeon 1982:221-239

Hong Kong

Baetis T₁

-"-

Hong Kong

Baetis T₂

-"-

Hong Kong

Baetis T₃

-"-

Hong Kong

Baetis T₄

Dudgeon 1984:55-129

Hong Kong

Baetis T₁

-"-

Hong Kong

Baetis T₂

-"-

Hong Kong

Baetis T₃

-"-

Hong Kong

Baetis T₄

-"-

Hong Kong

Baetis T₅

-"-

Hong Kong

Baetis L₅

-"-

Hong Kong

Baetis L₆

-"-

Hong Kong

Baetis L₇

-"-

Hong Kong

Baetis L₈

Landa & Soldan 1983

  Mongolia

Baetis sp.I

-"-

  Mongolia

Baetis sp.II

Furth 1983

Arabia

Baetis sp.1.6

-"-

Arabia

Baetis sp.1.56

-"-

Arabia

Baetis sp.

Wolda & Flowers 1985*

  Panama 

Baetis sp.

Bae 1985

 Korea

Baetis KUa

mutica [Ephemera] 

-"-

 Korea

Baetis KUb

-"-

 Korea

Baetis KUc

-"-

 Korea

Baetis KUd

Naranjo 1986:118-122*

 Cuba

Baetis sp.1

-'-

 Cuba

Baetis sp.2

-'-

 Cuba

Baetis sp.3

-'-

 Cuba

Baetis sp.4

Kobayashi 1987

-"-

 Japan

Baetis sp.D

Nigrobaetis sp.D

-"-

Baetis sp.E

parvipterus [Tenuibaetis] 

-"-

Japan

Baetis sp.G

Nigrobaetis sp.G

orientalis [Baetis atrebatinus]

-"-

 Japan

Baetis sp.I

Nigrobaetis sp.I

latus [Nigrobaetis]

-"-

 Japan

Baetis sp.L

bicaudatus [Baetis]  

-"-

 Japan

Baetis sp.N

Nigrobaetis sp.N

-"-

 Japan

Baetis sp.P

Nigrobaetis sp.P

apterus [Nigrobaetis]

ishigakiensis  [Nigrobaetis]

-"-

Japan

Baetis sp.Q

Nigrobaetis sp.Q

tricolor [Baetis]

Jamasaki 1987

Baetis sp.G: Kobayashi 1987

orientalis [Baetis atrebatinus]

Vincon & Thomas 1987*

  Pirenees

Baetis sp.A

Sivaramakrishnan & 
Venkataraman 1987

India

Baetis sp.A

-"-

India

Baetis sp.B

Salem & Ahmad 1987*

Baetis sp.

Joon & Bae 1988

Baetis n1a: Imanishi 1940

fuscata [Ephemera]  

-"-

Baetis KUa

mutica [Ephemera] 

Kobayashi 1989

Baetis sp.G: Kobayashi 1987

orientalis [Baetis atrebatinus]

-"-

Baetis sp.Q: Kobayashi 1987

tricolor [Baetis]

Arens 1989*

 mouth app.

Baetis sp.

Arens 1990*

 SEMmd

Baetis sp.

Flowers 1990*

Fuji

Baetis sp.gr.molawinensis

Flowers 1991*

 Panama 

Baetis sp.A

-"-

 Panama 

Baetis sp.B

-"-

 Panama 

Baetis sp.C

Soldan 1991*

Vietnam

Baetis sp.

-"-

Vietnam

Baetis sp.

-"-

Vietnam

Baetis sp.

McCafferty & Davis 1992

Baetis sp.1

punctiventris [Cloeon]

Gupta 1993

India

Baetis sp.A

-"-

India

Baetis sp.B

Novikova & Kluge 1994

FE Russia

Baetis (Nigrobaetis?) sp.

Bae & Andrikovics 1997*

 N.Korea

Baetis sp.1

pseudothermicus [Baetis]

-"-

 N.Korea

Baetis sp.2

silvaticus [Baetis]

Bae & Soldan 1997*

 N.Korea

Baetis sp.A

pseudothermicus [Baetis]

-"-

 N.Korea

Baetis sp.B

silvaticus [Baetis]

-"-

 N.Korea

Baetis sp.C

ursinus [Baetis]

Pardo & Campbell & Brittain 1998

Baetis sp.A

-"-

Baetis sp.B

-"-

Baetis sp.3

Nakamura & Kameyama 
& Katayama 2001

Japan

Baetis sp.E

Ishiwata & Kobayashi 2003

Baetis sp.G: Kobayashi 1987

orientalis [Baetis atrebatinus]

Mora & Boda & Csabai & 
Deak & Malnas & Csepes 2005

Baetis sp.

pentaphlebodes [Baetis]

Fujitani 2006

 Japan

Baetis sp.F

-"-

 Japan

Baetis sp.J

-"-

 Japan

Baetis sp.M

-"-

 Japan

Baetis sp.O

Hayashi & Nakano 2007

Baetis sp.1

-"-

Baetis sp.2

-"-

Baetis sp.3

-"-

Baetis sp.4

-"-

Baetis sp.5

Savolainen & Drotz & Hoffsten & Saura 2007

Baetis sp.n.

in gr. vernus

Willkomen 2008*

 pterothorax

Baetis sp.

Beketov 2008:289-299

  Siberia

Baetis sp.1

-"-

  Siberia

Baetis sp.2

Iwasaki & Kagaya & Miyamoto & Matsuda 2009

Baetis sp.H

Alhejoj & Salameh & Bandel 2014

Baetis sp.

pacis [Baetis] 

Kluge 2020:438-464.

  S.Africa

Protopatellata gen.sp.1

? Austrocloeon

? africanum [Cloeon]

-"-

  S.Africa

Protopatellata gen.sp.2

Kluge & Srinivasan & Sivaruban & Barathy & Isack 2023

Thailand

Baetis (Tenuibaetis) sp.

Baetis sp.

Baetis sp.

Baetis sp.

Baetis sp.

Baetis sp.

Baetis sp.