EPHEMEROPTERA OF THE WORLD

arb.sig. ABC

z   

Caenis/fg (= Coenis/fg)

f: Caenidae (orig. Coenidae) Newman 1853: 187

g: Caenis Stephens 1835: 60, typus C. macrura Stephens 1835 (design. Westwood 1840: Synopsis, p. 47)

Nomina circumscribentia: 


Caenoidea

Caenidae

Caeninae

Caenini

genus Caenis

taxa subordinated

binomina

 

Caenidae genn.spp.

Caenis spp.


Caenis/f1

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 252

 

Caenis/f1=Brachycercus/g1 = Caenotergaliae

 

Furcatergaliae

 

Kluge 2004

Caenis/f1=Brachycercus/g1 = Caenotergaliae

= Caenoidea: Edmunds & Traver 1954

Fimbriatotergaliae

Neoephemera/fg1 

Caenoptera

 

 

Caenis/f1

 

 

 

 

Caenis/f1=Brachycercus/g1 = Caenotergaliae

 

Fimbriatotergaliae

Neoephemera/fg1 

Caenoptera

 

 

 

 

 

 


Caenis/f2

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2000: 252

 

Caenis/f2=Brachycercus/g2 = Caenoptera

 

Caenotergaliae

 

Kluge 2004

Caenis/f2=Brachycercus/g2 = Caenoptera

= Brachycercidae: Lestage 1924

= Caenidae: Lestage 1942

= Caeninae: Edmunds & Traver 1954

= Brachycercus: Curtis 1834

= Caenis: Stephens 1835

Caenotergaliae

Brachycercus/f1=g3 

Caenis/f3=g1

 

 

 

 

 

 

 

Caenis/f2=Brachycercus/g2 = Caenoptera

 

Caenotergaliae

Brachycercus/f1=g3 

Caenis/f3=g1

 

 

 

 

 

 


Caenis/f3=g1

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 285

Caenis/f3=g1

 

Caenoptera

Madecocercus  

Caenis/f4=g2

 

 

 

 

 

 

 

Caenis/f3=g1

 

Caenoptera

 

 

 

 

 

 

 


Caenis/f4=g2

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 286

Caenis/f4=g2

= Caeninae: Malzacher 1997

Caenis/f3=g1

pm. Tillyardocaenis

Caenis/f5=g3

 

 

 

 

 

 

 

Caenis/f4=g2

 

Caenis/f3=g1

 

 

 

 

 

 

 


Caenis/f5=g3

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 287

Caenis/f5=g3

 

Caenis/f4=g2

pm. Caenis/f6=g4, 

Brasiliocaenis 

Clypeocaenis/g1

Caenopsella/g1

 

 

 

 

 

 

 

Caenis/f5=g3

 

Caenis/f4=g2

 

 

 

 

 

 

 


Caenis/f6=g4

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Kluge 2004: 287

pm. Caenis/f6=g4

 

Caenis/f5=g3

 

 

 

 

 

 

 

 

pm. Caenis/f6=g4

 

Caenis/f5=g3

 

 

 

 

 

 

 


informal taxa

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Eaton 1883-1888

section 7 of Caenis

 

Ephemeridae

group II, series III

Tricorythus

Lepthyphes

Caenis

 

 

 

 

 

 


superfamilia Caenoidea

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Spieth 1933

 

Caenoidea

 

 

 

Edmunds & Traver 1954

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Edmunds 1962

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Tshernova 1962

 

Caenoidea

 

 

Caenidae

Edmunds & Allen & Peters 1963

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Tshernova 1970

 

Caenoidea

 

Plectoptera

Caendae

Prosopistomatidae

Koss 1973

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Riek 1973

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Edmunds & Jensen & Berner 1976

 

Caenoidea

 

 

Neoephemeridae

Caenidae

McCafferty & Edmunds 1979

 

Caenoidea

 

Pannota

Neoephemeridae

Caenidae

Tshernova 1980

 

Caenoidea

 

 

Neoephemeridae

Caenidae

Prosopstomatidae

Edmunds 1982*

Caenoidea

 

Pannota

Neoephemeridae

Caenidae

Landa & Soldan 1985

 

Caenoidea

 

Pannota

Neoephemeridae

Baetiscidae

Caenidae

Prosopstomatidae

Hubard 1987

 

Caenoidea

 

Pannota

Neephemeridae

Caenidae

Kluge 1989

 

Caenoidea

 

Furcatergalia

Neoephemeridae

Caenidae

Peters & Campbell 1991

 

Caenoidea

 

Pannota

Neoephemeridae

Caenidae

Prosopistomatidae

Baetiscidae

McCafferty 1991

 

Caenoidea

 

Pannota

Caendae

Prosopistomatidae

Tricorythidae

Ephemerellidae

Kluge 1993

 

Caenoidea

 

Furcatergalia

 

McCafferty & Wang 2000*

Caenoidea

 

Pannota

Neoephemeridae

Caenidae

Jacobus & McCafferty 2006:81-90*

Caenoidea

 

Pannota

Neoephemeridae

Caenidae

 

 

Caenoidea

 

 

 

 

Caenis/f1=Brachycercus/g1

Caenis/f2=Brachycercus/g2

 

 

 

 

 

 

 

 

 


familia Caenidae (= Coenidae)

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Newman 1853

wings 2; caud.3

Coenidae fam.n. 

 

Ephemerina

 

Klapalek 1909: 14

 

Caenidae

 

Ephemerida

 

Ulmer 1920:97-144*

Caenidae

 

Baetoidea (subordo)

Caenis

Eurycaenis

Tricorythus

Tricorythodes

Leptohyphes

Leptohyphodes

Lestage 1924:316-352*

 

Caenidae

 

Baetoidea

 

Lestage 1924:61-62*

 

syn. Brachycercidae nom.n.

 

 

 

Eastham 1934*

tergalii

 

 

 

 

Ulmer 1939*

Caenidae

 

 

Caens

Tricorythus

et al.

Lestage 1942*

Caenidae (sens.restrict.)

 

 

Austrocaenis

Brachycercus

Caenodes

Caenis

Edmunds & Traver 1954

 

Caenidae

 

Caenoidea

Caeninae

Demoilin 1958

 

Caenidae

 

Ephemerelloidea

 

Tshernova 1959

 

Caenidae

 

 

 

Demoulin 1961

 

Caenidae

 

 

 

Edmunds 1962

 

Caenidae

 

Caenoidea

 

Edmunds & Allen & Peters 1963

 

Caenidae

 

Caenoidea

 

Riek 1970

 

Caenidae

 

Ephemeroidea

 

Tshernova 1970

 

Caenidae

 

Caenoidea

 

Landa 1973

 

Caenidae

 

 

 

Koss 1973

 

Caenidae

 

Caenoidea

 

Riek 1973

 

Caenidae

 

Caenoidea

 

Edmunds & Jensen & Berner 1976

Caenidae

 

Caenoidea

 

McCafferty & Edmunds 1979

 

Caenidae

 

Caenoidea

 

Tshernova 1980

 

Caenidae

 

Caenoidea

 

Edmunds 1982*

Caenidae

 

Caenoidea

 

Landa & Soldan 1985

 

Caenidae

 

Caenoidea

 

Tshernova & Kluge & 
Sinitshenkova & Belov 1986

Caenidae

 

 

 

Kluge 1989

 

Caenidae

 

Caenoidea

 

Provonsha 1990*

Caenidae

 

 

 

McCafferty 1991

 

Caenidae

 

Caenoidea

 

Malzacher 1991:VI.Conf.Eph.

genit.

Caenidae

 

 

 

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Caenidae

 

Furcatergalia

 

Malzacher 1997

 

Caenidae

 

 

Caeninae

Brachycercinae

Wang & McCafferty & Bae 1997*

phylog.

Caenidae

 

 

 

Kluge 1997:176-220

 

Caenidae

 

 

 

McCafferty & Wang 2000

Caenidae

 

Caenoidea

Caeninae

Brachycercinae

Madecocercinae

Jacobus & McCafferty 2006:81-90*

Caenidae

 

Caenoidea

Caeninae

Brachycercinae

Madecocercinae

 

 

Caenidae

 

 

 

 

Caenis/f1=Brachycercus/g1

Caenis/f2=Brachycercus/g2

 

 

 

 

 

 

 

 

 


subfamilia Caeninae (= Coeninae)

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Jacobson & Bianchi 1905

 

Caenidinae

 

Leptophlebiidae

Caenis

Tricorythus

Prosopistoma

Lameere 1917

 

Caeninae

 

Ephemeridae

Caenini

Ephemerellini

Lestage 1917

Coeninae

 

Baetidae

?Tricorythus (Coenis maxima)

Coenis

Handlirsch 1925

 

Caeninae

 

Ephemeridae

 

Needham & Traver & Hsu 1935

Caeninae

 

Baetidae

Caenis

Brachycercus

Tricorythodes

Edmunds & Traver 1954

 

Caeninae

 

Caenidae

Caenis

Brachycercus

Austrocaenis

Tasmanocaenis

Malzacher 1987

 

Caeninae

 

Caenidae

 

Malzacher 1997:VIII.Conf.Eph.

Caeninae

 

Caenidae

Caenis 

Tasmanocaenis

Brasiliocaenis

Clypeocaenis

Amercaenis

Barnardara

Caenopsella

Afrocaenis

Caenoculis

Wundacaenis

McCafferty & Wang 2000

Caeninae

 

Caenidae

 

Jacobus & McCafferty 2006:81-90

 

Caeninae

 

Caenidae

 

Malzacher & Staniczek 2006

Caeninae

 

Caenidae

 

 

 

Caeninae

 

Caenidae

 

 

Caenis/f1=Brachycercus/g1

Caenis/f2=Brachycercus/g2

Caenis/f3=g1

Caenis/f4=g2

Caenis/f5=g3

Caenis/f6=g4

 

 

 

 

 

 

 

 

 


tribus Caenini

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Banks 1900

 

Caenini

 

Ephemeridae

Caenis

Lameere 1917

 

Caenini

 

Ephemeridae-Caeninae

Caenis

Tricorythus

 

 

Caenini

 

 

 

 

Caenis/f1=Brachycercus/g1

Caenis/f2=Brachycercus/g2

Caenis/f3=g1

Caenis/f4=g2

Caenis/f5=g3

Caenis/f6=g4

 

 

 

 

 

 

 

 

 


genus Caenis (= Coenis)

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Stephens 1835*

Caenis gen.n.

= Brachycercus

Anisoptera=Ephemeridae

[subgen.]A.Caenis

[subgen.]B.Brachycercus

Pictet 1843-1845

Caenis

 

 

 

Brauer 1857 

Caenis

= Oxycypha syn.n. 

Ephemeridae

 

Walsh 1862*

Caenis

 

Ephemerina

 

Hagen 1863:1-35'

 
(unknown)

Caenis

= Brachycercus

= Oxycypha

Ephemeridae

 

Eaton 1868:82-91*

-

Caenis

= Oxycypha

= Brachycercus

 

 

Eaton 1871*

Caenis

= Oxycypha  

= Brachycercus  

= Macrocercus syn.n.

= Coeneus

 

 

Blackburn 1882:53-58.

Caenis

 

 

 

Eaton 1883-1888

Caenis

 

Ephemeridae

section 7 of Caenis

 

Schiller 1890

Caenis

 

 

 

Needham & Betten 1901*

Caenis

 

Ephemeridae-Baetinae

 

Needham 1905*

Caenis

 

Ephemeridae-Baetinae

 

Jacobson & Bianchi 1905

 

Caenis

 

Leptophlebiidae-Caenidinae

 

Lestage 1917

Coenis

 

Baetidae-Coeninae

 

Lestage 1919

Coenis

 

Baetidae-Coeninae

 

Ulmer 1920:97-144*

Caenis

 

Caenidae

 

Lestage 1924:61-62*

 

syn. Brachycercus

sensu auct = Ordella

 

 

McDunnough 1931

 

Caenis

 

 

 

Spieth 1933

phylog.

Caenis

 

Caenidae

 

Needham & Traver & Hsu 1935

Caenis

= Ordella  

Baetidae-Caeninae

 

Eastham 1936

 

Caenis

 

 

 

Ulmer 1939*

Caenis

 

Caenidae

 

Lestage 1942

Caenis

= Ordella

Caenidae

 

Traver 1944

Caenis

 

 

 

Grandi 1951

Caenis

 

 

 

Edmunds & Traver 1954

 

Caenis

 

Caenidae=Caeninae

 

Thew 1960

Caenis

 

 

 

Thibault 1970*

sex.char.

Caenis

 

 

 

Jacob 1974:93-97

 

Caenis

= Oxycypha  

= Ordella

 

 

Edmunds & Jensen & Berner 1976

Caenis

 

Caenidae

 

Demoulin 1981'

Caenis

 

Caenidae

 

Belfiore 1984*

 

Caenis

 

 

gr.macrura

gr.rhenicola

gr.pseudorivulorum

Roldon 1985*

Caenis

 

 

 

Kluge 1985: 1588

 

Caenis

= Caenomedea syn.n.  

= Caenodes syn.n.

 

 

Tshernova & Kluge & 
Sinitshenkova & Belov 1986

Caenis

 

Caenidae

 

Malzacher 1987

 

Caenis

 

Caenidae-Caeninae

 

Provonsha 1990*

Caenis

 

 

gr.diminuta

gr.hilaris

Studemann & Landolt & 
Sartori & Hefti & Tomka 1992

Caenis

 

Caenidae

 

McCafferty & Flowers & Waltz 1992*

biogeogr.

Caenis

 

Caenidae

 

Dominguez & Hubbard & Peters 1992*

Caenis

 

Caenidae

 

Malzacher 1993: 414

Caenis

= Caenomedea  

= Caenodes   

= Austrocaenis syn.n.

 

 

Kluge 1997:176-220

 

Caenis 

= Oxycypha 

= Ordella

Caenidae

 

Malzacher 1997:VIII.Conf.Eph.

 

Caenis

 

Caenidae-Caeninae

 

Dominguez & Molineri 
& Pescador &  Hubbard
& Nieto 2006

 

Caenis 

 

Caenidae

 

Malzacher 2009

Caenis

 

 

 

 

 

Caenis

 

 

 

 

Caenis/f1=Brachycercus/g1

Caenis/f2=Brachycercus/g2

Caenis/f3=g1

Caenis/f4=g2

Caenis/f5=g3

Caenis/f6=g4

 

 

 

 

 

 

 

 

 


subgenus  Caenis (= Coenis)

sources:

subjects:

valid name(s):

syn.:

higher taxon

subord.taxa:

Stephens 1835*

A. Caenis mihi

 

Caenis gen.n.

 

 

 

subgen. Caenis

 

 

 

 

Caenis/f3=g1

Caenis/f4=g2

Caenis/f5=g3

Caenis/f6=g4

 

 

 

 

 

 

 

 

 


taxa subordinated to Canis/f1=g.. & Caenoidea (> Caenoptera)

 Baetisca/fg [f:1900; g:1862] and subordinated taxa

 Ephemerella/fg [f:1909; g:1862] and subordinated taxa

 Neoephemera/fg [f:1935; g:1925] and subordinated taxa

 Prosopistoma/fg [f:1917; g:1833] and subordinated taxa

 Tricorythus/fg [f:1942; g:1868] and subordinated taxa


taxa subordinated to Canis/f2=g.. & Caenidae & Caeninae & Caenis (in Caenoptera)

Aenigmocaenis/g [g:2009]

Afrocaenis/g [g:1982]

Afrocercus/g [g:1987]

Alloretochus/g [g:2008]

Amercaenis/g [g:1985]

Austrocaenis/g [g:1932]

Barnardara/g [g:1995]

Brachoculis/fg [fg:2017]

Brachycercus/fg [f:1924; g:1834]

Brasiliocaenis/g [g:1975]

Caenoculis/fg [f:2008; g:1986]

Caenodes/g [g:1924]

Caenomedea/g [g:1960]

Caenopsella/g1 [g:1977

Callistellina/g [g:2004]

Callistina/g [2001] syn.obj. Callistellina

 Cercobrachys/fg [f:2008; g:1986] 

Clypeocaenis/g [g:1978]  

 Eurycaenis/g [g:1917] syn.obj. Brachycercus

 Insulibrachys/fg [f:2008; g:1986]

Irpacaenis/g [g:1999]

Kalimaenis/g [g:2013] 

Latineosus/fg [fg:2008]

 Madecocercus/fg [f:2000; g:1995]

Mandelara/g [g:2016]

Mccaffertya/g [2008] syn.obj. Callistellina

Niandancus/fg [f:2017; g:2009]

Ordella/g [g:1923] syn.obj. Caenis

Oriobrachys/g [fg:2008]

Oxycypha/g [g:1839]

Papuaenis/g [2016]

Provonshaka/g [g:1995]

Provonshara/g [g:2014]

Pseudocaenis/g [g:1978]

Sparbarus/fg [fg:2008]

Susperatus/g [g:2008]

Tasmanocoenis/g [g:1931]

Tigrocercus/g [g:2006]

Tillyardocaenis/g(1) [g:2004]

Trichocaenis/g [g:2009]

Wundacaenis/g [g:1993]


binomina

Caenidae genn.spp.

Caenis spp.