CLADOENDESIS OF EPHEMEROPTERA

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Hyrtanella/f1-Torleya/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Ephemerella/fg1
Tricorythus/fg1Ephemerella/fg2Timpanoga/fg1Ephemerella/fg3
- Hyrtanella/f1-Torleya/g1)

Nomen hierarchicum: Hyrtanella/f1=Torleya/g1 [f:1980; g:1917] (incl. Teloganopsis, Crinitella, Uracanthella, Amurella)

In circumscription fits:

— Torleya/g1: Kluge 2004: 312


References. Kluge 2004: * * * *


Autapomorphies of Hyrtanella/f1=Torleya/g1. Distinct autapomorphies are unknown. The features listed here, are characteristic for Torleya/g1, while can not serve as diagnostic to separate Torleya/fg1 it from Ephemerella/fg4.

(1) Maxillary palp is usually smaller than in majority of Ephemerella/fg3, often lost [see Ephemerella/fg1 (1)].

(2) Larval caudalii often (but not always) with whorls of heavy spine-like setae on posterior margins of segments, and often (but not always) without swimming setae.

Plesiomorphy of Hyrtanella/f1=Torleya/g1. Unlike Ephemerella/fg4, ventral lobe of tergalius VI is bifurcate (Kluge 2004: Fig.91:B), as that of tergalii III–V [see Ephemerella/fg3 (1)].

Size. Fore wing length 4–10 mm.

Distribution. Holarctic and Oriental Region.


The taxon Hyrtanella/f1=Torleya/g1 is divided into:

1. plesiomorphon Torleya/g2

2. Teloganopsis 

3. Hyrtanella/f2=Crinitella/g1 

4. Uracanthella 

5. Amurella 

Some species, known as larvae only, have uncertain systematic position