CLADOENDESIS OF EPHEMEROPTERA

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(1) Tergalii III are lost, thus bilamellate tergalii, whose structure is typical for Ephemerella/fg1 [see Ephemerella/fg1 (7)], are present on abdominal segments IV–VII only (Kluge 2004: Fig.91:I). Unique apomorphy (see Table).

Plesiomorphies and variable characters of Timpanoga/fg1. Unlike Ephemerella/fg3, tergalii I [stick-like – see Ephemerella/fg1 (13)] are usually retained, being attached to posterior part of the segment (Kluge 2004: Fig.91:G–H) (while in other mayflies with stick-like tergalii I – Vietnamella, Melanemerella/fg1, Caenotergaliae and Potamanthus/fg1 – they are attached anteriorly); in Timpanoga/fg4 tergalii I are transferred to extreme posterior margin of the segment; in Attenella/g1 delantala [Ephemerella]) tergalii I are lost. Paired tergal projections [see Ephemerella/fg1 (19)] are nearly always retained at least on some segments, rarely lost (in Dannella). Unlike Teloganopsis and Hyrtanella/f2=Crinitella/g1, hind wing without prominent costal projection and with Sc terminating near apex.