CLADOENDESIS OF EPHEMEROPTERA

ABC

z 

Rhithrogena/fg4

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata 
Rhithrogena/fg1
Epeorus/fg1Rhithrogena/fg2CinygmulaRhithrogena/fg3 - Rhithrogena/fg4)

Nomen hierarchicum: Rhithrogena/fg4 [f:1917; g:1881] (sine Epeiron, Himalogena, Sibirigena 

In circumscription fits:

Rhithrogena/fg4: Kluge 2004: 194

Non-typified taxa in Rhithrogena/fg4:

laevigata Tomka & Rash 1993

lobata Tomka & Rash 1993


Reference. Kluge 2004: * *


Characters of Rhithrogena/fg4 of unclear phylogenetic status.

(1) In imago and subimago 1st segment of fore tarsus is short, several times shorter than 2nd. The same in Sibirigena, unlike Himalogena and Epeiron/g1; the same in some other taxa (see Index of characters [2.2.76]).

(2) Each penis lobe is more or less tubular, with gonopore apically; sides near apex usually bear spines or denticles, including latero-dorsal [see Pentamerotarsata (5)] and medio-dorsal ones; titillators [ventral see Rhithrogena/fg3 (5)] are always well-developed (Sowa 1970: Fig.121; Tomka & Hasler 1978: Fig.14). The same in Himalogena, unlike Sibirigena and Epeiron).

Size. Fore wing length 517 mm. 

Distribution. Holarctic.


Nominal species in Rhithrogena/fg4:

adrianae Belfiore 1983 [Rhithrogena]  

allobrogica Sowa & Degrange 1987 [Rhithrogena]

alpestris Eaton 1885 [Rhithrogena]

alpicola Navás 1935 [Rhithrogena] syn.subj. alpestris [Rhithrogena]

austriaca Sowa & Weichselbaumer 1988 [Rhithrogena]

ayadi Dakki & Thomas 1986 [Rhithrogena]

 basalis [Baetis] nomen nudum   syn.subj. semicolorata [Baetis

bajkovae Sowa 1973 [Rhithrogena --/

beskidensis Alba-Tercedor & Sowa 1987 [Rhithrogena]

binotata Sinitshenkova 1982 [Rhithrogena] syn.subj. lepnevae [Rhithrogena]

bogoescui Sowa & Degrange 1987 [Rhithrogena]

braaschi Jacob 1974 [Rhithrogena]

brenneriana Klapálek 1905 [Rhithrogena] syn.subj. alpestris [Rhithrogena]

brodskyi Kustareva1976 [Rhithrogena]

brunneotincta McDunnough 1933 [Rhithrogena

bulgarica Braasch & Soldán & Sowa 1985 [Rhithrogena]

buresi Sowa 1973 [Rhithrogena]

carpatica Sowa 1971 [Rhithrogena picteti]  

carpatoalpina Klonowska & Olechowska & Sartori & Weichselbaumer 1987 [Rhithrogena]

castellana Navás 1927 [Rhithrogena]

caucasica Braasch 1979 [Rhithrogena] --/

cincta Navás 1921 [Rhithrogena]

circumtatrica Sowa & Soldán 1986 [Rhithrogena]

colmarsensis Sowa 1984 [Rhithrogena]

comitissa Navás 1933 [Rhithrogena] syn.subj. gorrizi [Rhithrogena]

corcontica Sowa & Soldán 1986 [Rhithrogena]

dagestanica Braasch 1979 [Rhithrogena]

daterrai Sowa 1984 [Rhithrogena]

decolorata Sinitshenkova 1973 [Rhithrogena]

degrangei Sowa 1969 [Rhithrogena]

delphinensis Sowa & Degrange 1987 [Rhithrogena]

diaphana Navás 1917 [Rhithrogena]

diensis Sowa & Degrange 1987 [Rhithrogena]

dorieri Sowa 1971 [Rhithrogena]

eatoni Esben-Petersen 1912 [Rhithrogena]

endenensis Metzler & Tomka & Zurwerra 1985 [Rhithrogena]

excisa Sinitshenkova 1979 [Rhithrogena]

expectata Braasch 1979 [Rhithrogena] --/

fasciata Traver 1933 [Rhithrogena]

ferruginea Navás 1905 [Rhithrogena]

fiorii Grandi 1953 [Rhithrogena]

fonticola Sowa & Degrange 1987 [Rhithrogena]

formosicola Matsumura 1931 [Rhithrogena] syn.subj. parvus [Ecdyonurus]

fradleyi Blair 1929 [Rhithrogena] syn.subj. germanica [Rhithrogena]

germanica Eaton 1885 [Rhithrogena]  

giudicellorum Thomas & Bouzidi 1986 [Rhithrogena]

goeldlini Sartori & Sowa 1988 [Rhithrogena]

gorganica Klapálek 1907 [Rhithrogena]

gorrizi Navás 1913 [Rhithrogena]

gratianopolitana Sowa & Degrange & Sartori 1986 [Rhithrogena]

grischuna Sartori & Oswald 1988 [Rhithrogena]

grisoculata Bogoescu 1933 [Rhithrogena] syn.subj. semicolorata [Baetis]

haarupi Esben-Petersen 1909 [Rhithrogena] syn.subj. germanica [Rhithrogena]

henschii Klapálek 1906 [Rhithrogena]

hercegovina Tanasijevic 1985 [Rhithrogena]

hercynia Landa 1969 [Rhithrogena]

hybrida Eaton 1885 [Rhithrogena]

ingalik Randolph & McCafferty 2005 [Rhithrogena]

insularis Esben-Petersen 1913 [Rhithrogena]

intermedia Zurwerra & Metzler & Tomka 1987 [Rhithrogena]

iranica Braasch 1983 [Rhithrogena]

iridina Kolenati 1859 [Baetis]

jacobi Braasch & Soldán 1988 [Rhithrogena]

jahorinensis Tanasievic 1985 [Rhithrogena]

japonica Ueno 1928 [Rhithrogena]

johannis Belfiore 1990 [Rhithrogena] syn.subj. siciliana [Rhithrogena]

joostiana Sowa & Zimmermann 1976 [Rhithrogena] --/

kimminsi Thomas 1970 [Rhithrogena]

klausnitzeriana Braasch 1979 [Rhithrogena]

klugei Tiunova 2010 [Rhithrogena]

kownackii Sowa & Zimmermann 1976 [Rhithrogena iridina]  

laciniosa Sinitshenkova 1979 [Rhithrogena]

landai Sowa & Soldán 1984 [Rhithrogena]

lepnevae Brodsky 1930 [Rhithrogena] --/

lisettae Baeuernfeind 2003 [Rhithrogena]

loyolaea Navás 1922 [Rhithrogena]

lucida Braasch 1979 [Rhithrogena]

marcosi Alba-Tercedor & Sowa 1987 [Rhithrogena]

mariae Vitte 1991 [Rhithrogena]

mariaedominicae Sowa & Degrange 1987 [Rhithrogena]

marinkovici Tanasijevic 1985 [Rhithrogena]

monserrati Alba-Tercedor & Sowa 1986 [Rhithrogena]

neretvana Tanasijevic 1985 [Rhithrogena]

nivata Eaton 1871 [Heptagenia]

nuragica Belfiore 1987 [Rhithrogena]

orientalis You 1990 [Rhithrogena]

oscensis Navás 1927 [Rhithrogena]

ourika Thomas & Mohati 1985 [Rhithrogena]

parvus Ulmer 1912 [Ecdyonurus]

picteti Sowa 1971 [Rhithrogena]

piechockii Braasch 1977 [Rhithrogena]

podhalensis Sowa & Soldán 1986 [Rhithrogena]

pontica Sowa & Soldán & Kazanci 1986 [Rhithrogena]

potamalis Braasch 1979 [Rhithrogena]

puthzi Sowa 1984 [Rhithrogena]

puytoraci Sowa & Degrange 1987 [Rhithrogena]

quadrinotata Sinitshenkova 1982 [Rhithrogena] syn.subj. bajkovae [Rhithrogena

reatina Sowa & Belfiore 1984 [Rhithrogena]

robusta Dodds 1923 [Rhithrogena]

rolandi Weichselbaumer 1995 [Rhithrogena]

rubicunda Traver 1937 [Rhithrogena]

ryszardi Thomas & Vitte & Soldán 1987 [Rhithrogena]

sangangensis You 1990 [Rhithrogena]

sartorii Zrelli & Boumaiza (in Zrelli & Sartori & Bejaoui & Boumaiza) 2011 [Rhithrogena]

savoiensis Alba-Tercedor & Sowa 1987 [Rhithrogena] /

semicolorata Curtis 1834 [Baetis] typus nominis Rhithrogena /

semitincta Pictet 1843 [Baetis] syn.subj. semicolorata [Baetis]

sibillina Metzler & Tomka & Zurwerra 1985 [Rhithrogena]

siciliana Braasch 1989 [Rhithrogena]  

soteria Navás 1917 [Rhithrogena]

sowai Puthz 1972 [Rhithrogena]

strenua Thomas (in Thomas & Gazagnes) 1982 [Rhithrogena]

sublineata Kazanci & Braasch 1988 [Rhithrogena]

tatrica Zelinka 1953 [Rhithrogena]

taurisca Bauernfeind 1992 [Rhithrogena]

teberdensis Zimmermann 1977 [Rhithrogena] --/

theischingeri Braasch 1981 [Rhithrogena]

thomasi Alba-Tercedor & Sowa 1986 [Rhithrogena]

thracica Sowa & Soldán & Braasch 1988 [Rhithrogena]

trispina Zhou & Zheng 2000 [Rhithrogena]

umbrosa Braasch 1979 [Rhithrogena]

unicolor Tshernova 1952 [Rhithrogena] syn.subj. lepnevae [Rhithrogena]

ussingi Esben-Petersen 1910 [Rhithrogena] syn.subj. germanica [Rhithrogena]

vaillanti Sowa & Degrange 1987 [Rhithrogena]

vulpecula Klapálek 1905 [Rhithrogena] syn.subj. semicolorata [Baetis]

wolosatkae Klonowska 1987 [Rhithrogena]

zelinkai Sowa & Soldán 1984 [Rhithrogena]

zernyi Bauernfeind 1991 [Rhithrogena]

zhiltzovae Sinitshenkova 1979 [Rhithrogena]


See also:

Rhithrogena/fg3 INCERTAE SEDIS

Radulapalpata INCERTAE SEDIS

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