CLADOENDESIS OF EPHEMEROPTERA

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Raptoheptagenia

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Branchitergaliae
Heptagennota Pentamerotarsata Radulapalpata - Raptoheptagenia)

Nomen hierarchicum: Raptoheptagenia/g(1) [g:1987]

In circumscription matches:

gen. Raptoheptagenia Whiting & Lehmkuhl 1987a: 405

gen. Anepeorus: Burks 1953: 198 (partim: larva, non imago)

subfam. Anepeorinae Edmunds 1962:10 (larva, non imago)

Raptoheptagenia/g(1): Kluge 2004: 188


References. Burks 1953 (Anepeorus): ; Edmunds & Allen & Peters 1963 (Anepeorinae): ; Edmunds & Jensen & Berner 1976 (Anepeorinae): ; McCafferty & Provonsha 1986 (Anepeorus): ; Whiting & Lehmkuhl 1987a: ; Kluge 1993: ; 2004:


Autapomorphies of Raptoheptagenia.

(1) Mouth apparatus is strongly modified and specialized for carnivorism (Kluge 2004: Fig.60:AE): Labial palps, which in all other Radulapalpata (including carnivorous species) are specialized as scraping [see Radulapalpata (1)], in Raptoheptagenia have lost their scraping specialization and specialized as grasping: 2nd+3rd segment is sclerotized, pointed, thinned and curved in a form of hook, its setae are nearly completely reduced. Maxilla is biting, with enlarged canines and dentisetae, its apical margin is oblique, without typical for Pentamerotarsata pectinate setae [see Radulapalpata (3)] and without filtering setae. Hypopharynx is shorter and wider, and superlinguae [see (3)] are smaller than in other Radulapalpata; mandibles have shape typical for specialized predators: shortened perpendicular to axis of articulation, mola is tooth-like, without grater (McCafferty & Provonsha 1986: Fig.7,13).

(2) Tergalii are attached to ventral side of abdomen. Non-unique apomorphy (see Index of characters [1.3.22]), but not found in other Heptagennota.

Characters of Raptoheptagenia of unclear phylogenetic status. (3) Superlinguae [diminished see (1)] are somewhat curved laterally (McCafferty & Provonsha 1986: Fig.13). Probably, a common feature with Heptagenia/f5=g4.

(4) Outer margin of larval femur [flattened see Heptagennota (1)] bear long thin hairs and sparse stout pointed setae (Kluge 2004: Fig.60:F). Probably, a common feature with Heptagenia/f6=g5, while other Radulapalpata have a regular dense row of long stout setae.

(5) Larval caudalii lack whorls of stout setae on hind margins of segments. Among Radulapalpata the same in Cinygma and Rhithrogena/fg1, while in Heptagenia/f5=g4 whorls of stout setae are present.

(6) In imago and subimago 1st tarsal segment is shortened, on fore tarsus of male several times shorter than 2nd segment. Non-unique apomorphy (see Index of characters [2.2.76]); among Pentamerotarsata the same in Heptagenia/f8=g7, Dacnogenia and selected species of Kageronia, Stenonema/g1, Ecdyonurus/fg1, Rhithrogena/fg3.

Size. Fore wing length 911 mm. 

Distribution. Nearctic.


Nominal species in Raptoheptagenia/g(1): 1 species

cruentata Walsh 1863 [Heptagenia] typus nominis Raptoheptagenia

reversalis McDunnough 1924 [Heptagenia] syn.subj. cruentata [Heptagenia]


See also:

Radulapalpata INCERTAE SEDIS

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