CLADOENDESIS OF EPHEMEROPTERA

ABC

z 

† Plesiomorphon Permoplectoptera (or Protereisma/f1=Phtharthus/g1)

(Panephemeroptera Euephemeroptera - pm.Permoplectoptera)

Nomen hierarchicum: Protereisma/f1=Phtharthus/g1 [f:1917; g:1904]

Nomina circumscribentia:

— Protereismephemeridae Sellards 1907: 345

— Permoplectoptera Tillyard 1932: 117

In circumscription fits:

— fam. Protereismephemeridae: Sellards 1907: 345

— fam. Protereismidae: Lameere 1917: 45

— superfam. Protereismatoidea: Demoulin 1958: 6

— subordo Permoplectoptera: Tillyard 1932: 117

— Permoplectoptera, or Protereisma/f1=Phtharthus/g1: Kluge 2000: 243

Nominal taxa included:

Protereisma/fg [f:1917; g:1907] (typus P. permianum)

Protechma/g [g:1907] (typus P. acuminatum)

Prodromites/g [g:1924] (typus Prodromus rectus) = Prodromus/g [g:1907] nom.praeocc.

Bantisca/g [g:1907] (typus B. elongata )

Rekter/g [g:1907] (typus R. arcuatus) = Recter/g [g:1909]

Pinctodia/g [g:1907] (typus P. curta )

Mecus/g [g:1909] (typus Scopus gracilis) = Scopus/g [g:1907] nom.praeocc.

Esca/g [g:1909] (typus Therates planus)

Kukalova/fg [fg:1970] (typus K. americana)

Misthodotes/fg [f:1932; g:1909] (typus Dromeus obtusus) = Dromeus/g [g:1907] nom.praeocc.

Eudoter/fg [f:1954; g:1936] (typus E. delicatulus)

Palingeniopsis/fg [f:1938; g:1932] (typus P. praecox)

Loxophlebia/g [g:1928] (typus L. apicalis)

Mesephemera/fg [f:1917; g:1906] (typus Ephemera procera)

Jarmila/fg [fg:1970] (typus J. elongata)

Oboriphlebia/fg [fg:1980] (typus Kukalova moravica)

Mesoplectopteron/fg [f:1955; g:1918] (typus M. longipes)

Cretereisma/g [g:2007] (typus C. antiqua)

Alexandrinia/g [2012] (typus A. gigantea)

Ponalex [g:2012] (typus P. maximus)


References. Sellards 1907: ; – Tillyard 1932: ; – Carpenter 1933: ; – Kukalova 1968: ; – Tshernova 1970: ; – Carpenter 1979: .


Plesiomorphies of Permoplectoptera (unlike Euplectoptera). Wings are homonomous – i.e. hind wings have nearly the same size and venation as fore wings (Kluge 2004: Fig.14:B). At least in some representatives tergalii are present not only on abdominal segments I–VII, but on abdominal segments VIII–IX as well (Kluge 2004: Fig.14:D). For larva of americana [Kukalova], are described segmented tarsus and two claws, that resembles adult structure and differs from larval Euplectoptera (Kukalova 1968).

Size. Fore wing length 6–32 mm.

Age. From Early Permian to Early Cretaceous.


Nominal species in Permoplectoptera:

 

with Protereisma-type wing venation:

      larva with well-preserved venation of protoptera:

americana Demoulin 1970 [Kukalova]

      adults:

acuminatum Sellards 1907 [Protechma] 

arcuatus Sellards 1907 [Rekter]

curta Sellards 1907 [Pinctodia

directa Sinitshenkova & Vassilenko 2012 [Alexandrinia]

directum Carpenter 1979 [Protereisma

elongata Sellards 1907 [Bantisca — typus nominis Bantisca

extensis Sellards 1907 [Rekter] 

gigantea Sinitshenkova & Vassilenko 2012 [Alexandrinia] — typus nominis Alexandrinia

gracilis Sellards 1907 [Scopus — typus nominis Mecus

insigne Tillyard 1932 [Protereisma

latum Sellards 1907 [Protereisma

??? maximus Sinitshenkova & Aristov 2012 [Ponalex] — typus nominis Ponalex 

minus Sellards 1907 [Protereisma

permianum Sellards 1907 [Protereisma] 

planus Sellards 1907 [Terates]  — typus nominis Esca

rectus Sellards 1907 [Prodromus] 

sellardsi Tillyard 1932 [Protereisma

 

with Misthodotes-type wing venation:

biguttatus Tillyard 1932 [Misthodotes]

delicatulus Tillyard 1936 [Eudoter]  — typus nominis Eudoter

edmundsi Carpenter 1979 [Misthodotes]

obtusus Sellards 1907 [Dromeus]

ovalis Tillyard 1932 [Misthodotes]

sharovi Tshernova 1965 [Misthodotes]

stapfi Kinzelbach & Lutz 1984 [Misthodotes]

tshernovae Sinitshenkova & Vassilenko 2012 [Misthodotes]

zalesskyi Tshernova 1965 [Misthodotes]

 

with Palingeniopsis-type wing venation:

praecox Martynov 1932 [Palingeniopsis]

 

with Cretereisma-type wing venation:

 antiqua Willmann 2007 [Cretereisma] — typus nominis Cretereisma

 schwickertorum Willmann 2007 [Cretereisma]

 

adults with unknown wing venation:

apicalis Martynov 1928 [Loxophlebia]

lithophilus Germar 1842 [Tineites]

palaeon Weyenbergh 1874 [Anomalon]

prisca Germar 1839 [Sciaria]

procera Hagen 1862 [Ephemera] — typus nominis Mesephemera

rossenrayensis Guthorl 1967 [Protereisma]

speciosa Oppenheim 1888 [Ephemera] (nom. praeocc.)

uralicum Zalessky 1947 [Protereisma]

weyenberghi Handlirsch 1906 [Mesephemera]

 

larvae with unknown wing venation:

elongata Demoulin 1970 [Jarmila]  — typus nominis Jarmila

longipes Handlirsch 1918 [Mesoplectopteron] 

moravica Demoulin 1970 [Kukalova] — typus nominis Oboriphlebia

netshaevi Handlirsch 1904 [Phtharthus] — syn.subj. rossicus [Phtharthus]

quarta Hubbard & Kukalova-Peck 1980 [Oboriphlebia]

quinta Hubbard & Kukalova-Peck 1980 [Oboriphlebia]

rossicus Handlirsch 1904 [Phtharthus]

tertia Hubbard & Kukalova-Peck 1980 [Oboriphlebia]

sp.: larval paratype of sharovi Tshernova 1965 [Misthodotes]

__