CLADOENDESIS OF EPHEMEROPTERA

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Characters of Metreletus of unclear phylogenetic status.

(1) In cubital field of fore wing, veins, which go from CuA to wing margin [see Anteritorna (1)], are more or less substituted by one or two intercalaries (Demoulin 1951: Fig.1a-e,11a-d; 1952c: Fig.4; Ujhelyi 1960: Fig.5; Kluge 2007: Fig.4). Non-unique apomorphy (see Index of characters [2.2.52]).

(2) Larval claw on its inner side, besides irregular small denticles near base, has one row of greater denticles (Demoulin 1951: Fig.6b-d) (this character is known for balcanicus [M.] only, while leg structure of micus [A.] is unknown). The same in some Bidentiseta and others (see Index of characters [1.2.21]); probably non-unique autapomorphy.

(3) Imaginal penis has ventral plates [see Ameletus/fg1 (6)] very long and blunt (this is reliably known for balcanicus [Metretopus] only); on larval protopenis, each lobe has a prominent ventral plate separated by incision from lateral lobe (Kluge 2007: Fig.1-2) (unlike larval Ameletus, whose ventral plates are not separated from lateral lobes – Kluge 2007: Fig.10, Fig.21).

(4) Labrum has a deep apical incision (Kluge 2007: Fig.3) (unlike Ameletus, whose labrum has a shallow apical incision).

Plesiomorphy of Metreletus (unlike Ameletus/fg2). Glossae are not truncate, apically-ventrally with numerous irregular pointed setae (Kluge 2004: Fig.20:D) (this character is known for balcanicus [M.] only, while labium structure of micus [A.] is unknown).