CLADOENDESIS OF EPHEMEROPTERA

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Madecocercus

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Fimbriatotergaliae Caenotergaliae Caenoptera Caenis/f3=g1 - Madecocercus)

Nomen hierarchicum: Madecocercus/g(1) [f:2000; g:1995]

In circumscription fits:

gen. Madecocercus Malzacher 1995: 2

gen. Provonshaka McCafferty & Wang 1995: 179

subfam. Madecocercinae McCafferty & Wang 2000

Madecocercus/g(1): Kluge 2004: 286

Synonym: subjective synonym of  Madecocercus/g (typus tauroides [M.]) is Provonshaka/g [g:1995] (typus thomasorum [P.]) (subjective synonymy: tauroides [M.] = thomasorum [P.])


References. Malzacher 1995: ; McCafferty & Wang 1995: ; Elouard & Sartori 2001: ; Kluge 2004:


Autapomorphies of Madecocercus.

(1) Maxillary palp is lost. Non-unique apomorphy, the same in some Ephemerella/fg1 (see Index of characters [1.1.41]).

(2) Marginal processes of tergalii IV-VI are non-branched, thus only tergalius III retains branched processes [see Caenotergaliae (1) and Caenoptera (4)]. On all tergalii III-VI on proximal-anal margin processes are substituted by setae of similar size (Kluge 2004: Fig.85:CD).

(3) Larval body has peculiar shape: flattened and widened, femora strongly widened, with regular rows of long setae on outer and inner margins; dorsal surface of body and legs is warty, on fore femur warts form a transverse row (which originally was confused with a setal row in Pantricorythi).

Plesiomorphies of Madecocercus. Prosternum is wide and rectangular; metanotum without transverse crest, flat, with scutellum distinctly projected posteriorly (Malzacher 1995: Fig.1c,d) (the same in Brachycercus/f1=g3, unlike Caenis/f4=g2). Gonostylus with a longitudinal groove [see Caenoptera (5)].

Size. Larval body length 56 mm (see Caenoptera). 

Distribution. Madagascar.


Nominal species in Madecocercus/g(1): 1 species

bibulus Malzacher (in Malzacher & Staniczek) 2006 [Madecocercus]

tauroides Malzacher 1995 [Madecocercus]

thomasorum McCafferty & Wang 1995 [Provonshaka] typus nominis Provonshaka syn.subj. tauroides [Madecocercus]


See also:

Caenoptera INCERTAE SEDIS

Furcatergaliae INCERTAE SEDIS

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