CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Leptohyphes/fg2
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Furcatergaliae
Ephemerella/fg1
Pantricorythi
Tricoryptera
Leptohyphes/fg1
- Leptohyphes/fg2)
Nomen hierarchicum:
Leptohyphes/fg2 [f:1954; g:1882]
(sine Allenhyphes, Tricorythodes, Tricorythopsis)
In circumscription fits:
— gen. Leptohyphes: Kluge 1992c: 6
— Leptohyphes/fg2: Kluge 2004: 335
Synonym:
subjective synonyms of Leptohyphes/fg (typus eximius [L.])
is Bruchella/g [g:1920]
(typus nigra [B.]) (subjective synonymy:
eximius [L.] = nigra [B.])
References. Kluge 1992c: 
; – Hofmann & Sartori & Tomas 1999:
; – Baumgardner &
McCafferty 2000 (L. zalope): ; – Wiersema & McCafferty 2000:
; – Molineri 2003a:
; – Kluge 2004: *
* *
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Autapomorphies of Leptohyphes/fg2. (1) Costal flap of ventral lobe of gill operculum [see Leptohyphes/fg1 (1)] is reduced to a spine-like process reaching base of the next tergalius only (Kluge 2004: Fig.101:D–E). (2) In imago and subimago [shortly-moulting – see Tricoryptera (7)] lateropostnotal crest is either lost, or sharply curved toward anterior margin of lateropostnotum (Kluge 2004: Fig.102:B). In other Ephemerella/fg1 lateropostnotal crest is more straight and looks as a continuation of infrascutellar-lateropostnotal arm (Kluge 2004: Fig.97:D, 104:B). Plesiomorphies of Leptohyphes/fg2. Unlike Allenhyphes and Tricorythodes, ventral lobe of tergalii III-V retains processes in a form of one or several lamellae arising ventrally near base [see Leptohyphes/fg1 (1)] (Kluge 2004: Fig.101:F–H). Unlike Haplohyphes, mouth apparatus is non-specialized, maxilla [see Ephemerella/fg1 (16)] with 3 canines and 2 dentisetae [see Bidentiseta (1)] (Kluge 1992c: Fig.2). In imago and subimago: Scuto-lateropostnotal muscle of metathorax is not so large as in Tricorythodes [while transverse interscutal suture is present – see Leptohyphes/fg1 (4)]; correspondingly, posterior scutal protuberances do not convergent posteriorly (can be divergent or parallel – Kluge 2004: Fig.102:A), lateropostnotum [see (2)] is not fused with epimeron, and is separated by epimeron from coxal conjunctiva (Kluge 2004: Fig.102:B). Lateroparapsidal suture somewhat retains curvation initial for Ephemerella/fg1 [see Leptohyphes/fg1 (4)], merging with transverse interscutal suture laterad of medioparapsidal suture (Kluge 2004: Fig.102:A). Hind wings [of peculiar structure – see Leptohyphes/fg1 (2)] are present at least in male; in female they are either present, or lost; metathoracic wing muscles S.ESm and MTm are retained even in female which have no hind wings (Kluge 2004: Fig.102:C–D). 1st tarsal segment is short and fused with tibia. Gonostylus is 3-segmented, i.e. with distal segment [short – see Pantricorythi (2)] distinguishable at least in imago. |
Size. Fore wing length 4–8 mm (see Tricoryptera).
Distribution. Neotropical and south of Nearctic Regions.
Nominal species in Leptohyphes/fg2:
airuoca Nascimento & Molineri & Salles 2014 [Leptohyphes]
albipennis Molineri & Zuniga 2006 [Leptohyphes]
albus Mayo 1968 [Leptohyphes]
— syn.subj. tacajalo [Leptohyphes]
apache Allen
1967 [Leptohyphes]
bruchi Navás 1913 [Leptohyphes]
— syn.subj. eximius
[Leptohyphes]
carinus Allen 1973 [Leptohyphes]
coconuco Molineri & Zuniga 2006 [Leptohyphes]
comatus Allen 1967 [Leptohyphes]
cornutus
Allen 1967 [Leptohyphes]
echinatus Allen
& Roback 1969 [Leptohyphes]
— syn.subj. setosus [Leptohyphes]
ecuador Mayo 1968 [Leptohyphes]
eximius
Eaton 1882 [Leptohyphes]
/
,
//
/
ferruginus Allen & Brusca 1973 [Leptohyphes]
guadeloupensis
Hofmann & Sartori (in
Hofmann & Sartori & Thomas) 1999 [Leptohyphes]
hirsutus
Allen & Roback 1969 [Leptohyphes]
hispidus
Allen & Brusca 1973 [Leptohyphes]
— syn.subj. apache [Leptohyphes]
illiesi Allen 1967 [Leptohyphes]
invictus
Allen 1973 [Leptohyphes]
jodiannae Allen 1967 [Leptohyphes]
kukenan Molineri & Derka 2020 [Leptohyphes]
liniti
Wang & Sites & McCafferty 1998 [Leptohyphes]
?-/
lumas Allen & Brusca 1973 [Leptohyphes]
— syn.subj.
apache [Leptohyphes]
maculatus Allen 1967
[Leptohyphes]
mandibulus Baumgardner 2007 [Leptohyphes]
mollipes Needham & Murphy 1924 [Leptohyphes]
myllonotus Allen & Roback 1969 [Leptohyphes]
nebulosus Nascimento & Molineri & Salles 2014 [Leptohyphes]
nigra Navás 1920 [Bruchella]
— syn.subj. eximius
[Leptohyphes]
nigripennis Molineri & Zuniga 2006 [Leptohyphes]
nigripunctum
Traver 1943 [Leptohyphes]
pereirae Da-Silva 1993 [Leptohyphes]
— syn.subj. plaumanni [Leptohyphes]
peterseni Ulmer
1920 [Leptohyphes]
petersi
Allen 1967 [Leptohyphes]
piraticus Allen 1978 [Leptohyphes]
— syn.subj. ferruginus
[Leptohyphes]
plaumanni Allen 1967 [Leptohyphes]
populus Allen 1973 [Leptohyphes]
setosus Allen 1967 [Leptohyphes]
succinus Allen 1978 [Leptohyphes]
— syn.subj. apache
[Leptohyphes]
tacajalo Mayo 1968 [Leptohyphes]
tuberculatus Allen 1967 [Leptohyphes]
zalope Traver 1958 [Leptohyphes]
,
zelus Allen 1978 [Leptohyphes]
— syn.subj. zalope [Leptohyphes]
See also:
Leptohyphes/fg1 INCERTAE SEDIS
Furcatergaliae INCERTAE SEDIS
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