CLADOENDESIS OF EPHEMEROPTERA

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Ichthybotus

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Fimbriatotergaliae Fossoriae - Ichthybotus)

Nomen hierarchicum: Ichthybotus/fg(1) [f:1957; g:1899]

In circumscription fits:

gen. Ichthybotus Eaton 1899: 285

subfam. Ichthybotinae: Koss & Edmunds 1974: 295

fam. Ichthybotidae Demoulin 1957a: 336

Ichthybotus/fg1: Kluge 2000: 252


References. Eaton 1899: ; Tillyard 1923: ; Phillips 1930: ; 1931: ; Koss & Edmunds 1974: ; Kluge 2004: .


Characters of Ichthybotus of unclear phylogenetic status.

(1) Mandibular tusks [see Fimbriatotergaliae (8)] are curved medially (probably plesiomorphy, as the same in Potamanthus/fg1, Euthyplocia/fg1, Polymitarcys/f1=Ephoron/g2); dorsal and lateral sides of tusks with numerous tubercles (the same in Polymitarcys/f2=Ephoron/g3).

(2) Apex of larval fore tibia projects posteriad of tarsus base and forms two projections outer-apical and inner-apical ones, with incision between them. Similar in Hexagenia/fg1 and Palingenia/f2=g1 (Kluge 2004: Fig.74:DE); possibly, symplesiomorphy [see Fossoriae (1)].

(3) Tergalius I [see Fimbriatotergaliae (6)] is vestigial, bilamellate, without processes. The same in majority of Fossoriae (except for Behningia/fg1) and in Euthyplocia/fg1; possibly, symplesiomorphy.

(4) Imaginal and subimaginal paracercus of male is vestigial, while paracercus of female is subequal to cerci. Non-unique apomorphy (see Index of characters [2.3.22]); among Fossoriae the same in Cryptoprosternata.

Plesiomorphies of Ichthybotus. In larva: Maxilla with 3 canines and 2 dentisetae [see Bidentiseta (1)]. Maxillary palp is 3-segmented; labial palp is 3-segmented (unlike Cryptoprosternata). Larval (and adult) patella-tibial suture is developed on middle and hind legs only. Unlike Ephemera/fg9, all tergalii are attached to posterior part of segment.

In imago and subimago: Mesonotal suture is moderately stretched posteriorly, and subimaginal mesonotal cuticle has pigmented areas of characteristic shape [see Fimbriatotergaliae (2)]. Furcasternal protuberances are contiguous (unlike Campsurus/fg1, Caenotergaliae and some others see Index of characters [2.2.23]). In cubital field of fore wing several (34) branched veins go from CuA to basitornal margin [see Anteritorna (1)]. Claws of all legs (including fore legs of male) are ephemeropteroid. Gonostylus is 4-segmented (i.e. with expressed 1st segment and with 2 distal segments).

Size. Fore wing length 1522 mm. 

Distribution. New Zealand.


Nominal species in Ichthybotus/fg(1):

bicolor Tillyard 1923 [Ichthybotus]

hudsoni McLachlan 1894 [Ephemera] typus nominis Ichthybotus ,/,/

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