CLADOENDESIS OF EPHEMEROPTERA
Ephemerella/fg1 Pantricorythi Melanemerella/f1=Teloganodes/g1 - pm. Ephemerellina/g1)
Nomen hierarchicum: Ephemerellina/g1 [g:1924] (incl. Indoganodes, Lithogloea, Lestagella, Nadinetella)
In circumscription fits:
Nominal taxa included:
Nadinetella/g [g:1998] = Nadinella/g [g:1997] nom.praeocc.
? Manohyphella/g [g:1973] — see Ephemerella/fg1 incertae sedis
References. Kluge 2004: * ' *
Plesiomorphies of Ephemerellina/g1. In imago and subimago (unknown for jobini [Indoganodes], brincki Demoulin 1970 [Ephemerellina]): Eyes of male are large (unlike some Tricoryptera and others). Imaginal and subimaginal mesonotum (examined for harrisoni [L.] only) has structure primitive for Ephemerella/fg1 [see Ephemerella/fg1 (9) and (10)]: mesonotal suture is distinct, lateroparapsidal suture is curved laterally; subimaginal lateral pigmented area of mesonotum is bordered by relief line; scutellum is not modified, infrascutellum is present (as in Ephemerella/fg2 and Vietnamella, unlike Teloganodes/f1=g3, Teloganella and Tricoryptera). Cubital field of fore wing with vestiges of 2 bifurcate veins [see Ephemerella/fg1 (15)] variably developed, sometimes reduced to 1–4 intercalaries. Hind wing is well-developed, as long as 0.2–0.25 of fore wing length, with Sc reaching wing apex, RS branching, and veins behind MP1 developed (unlike Teloganodes/f1=g3, Teloganella and Tricoryptera).
Variable characters of Ephemerellina/g1. Larval head either has normal shape, or (in penicillata [L.]) is flattened with frons forming a wide flat shield, which is margined by a row of long setae and overlaps clypeus, labrum and mandibles; in this case, mandibles are strongly shortened (Demoulin 1970e: Fig.15a-d) (similar to Dicercomyzon, some Teloganodes/f1=g3 and others — see Index of characters [1.1.4]).
Each larval claws can bear 1 row of denticles [see Ephemerella/fg1 (18)], or 2 equal rows (in crassi [E.] and brincki [E.], united in Nadinetella) (the same in some other taxa — see Index of characters [1.2.21]).
Projections of posterior margin of larval abdominal terga [see Ephemerella/fg1 (19)] can be either vary individually from paired to unpaired (in crassi [E.] and brincki [E.]), or are constantly unpaired (in barnardi [E.], harrisoni [L.], animosa [M.], sphyxia [M.]), or absent (in penicillata [L.]) (Demoulin 1970e: Fig.12–15; McCafferty & Wang 1997: Fig.4–7, 74–76).
Number of tergalii is variable (see Table): Tergalii I [stick-like – see Ephemerella/fg1 (13)] are either present (in crassi [E.], penicillata [L.], harrisoni [L.], animosa [M.], sphyxia [M.]), or lost (in jobini [Indoganodes], barnardi [E.] and brincki [E.]); lamellate tergalii can be developed either up to pair VI (in jobini [Indoganodes], harrisoni [L.] and barnardi [E.]), or up to pair V (in brincki [E.] and crassi [E.]), or up to pair IV only (in penicillata [L.]).
Size. Fore wing length 5–12 mm.
Distribution. Afrotropical and Nearctic Regions.
Nominal species in Ephemerellina/g1:
barnardi Lestage 1924 [Ephemerellina] — typus nominis Ephemerellina
brincki Demoulin 1970 [Ephemerellina (Ephemerellina)]
crassi Allen & Edmunds 1963 [Ephemerellina] — typus nominis Nadinetella
harrisoni Barnard 1932 [Lithogloea] — typus nominis Lithogloea --/
jobini Selvakumar & Sivaramakrishnan & Jacobus (in Selvakumar & Sivaramakrishnan & Jacobus & Janarthyanan & Arumugam) 2014 [Indoganodes] — typus nominis Indoganodes
penicillata Barnard 1940 [Lithogloea] — typus nominis Lestagella /
Presumably here belong also Madagascar species:
animosa McCafferty & Benstead 2002 [Manohyphella]
sphyxia McCafferty & Benstead 2002 [Manohyphella]