CLADOENDESIS OF EPHEMEROPTERA
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CLADOENDESIS OF EPHEMEROPTERA |
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Cinygmula/g1
(Panephemeroptera
Euephemeroptera
Euplectoptera
Anteritorna
Bidentiseta
Branchitergaliae
Heptagennota
Pentamerotarsata
Radulapalpata
Rhithrogena/fg1Epeorus/fg1Rhithrogena/fg2
- Cinygmula)
Nomen hierarchicum:
Cinygmula/g1
[g:1933] (incl. Ororotsia)
In circumscription fits:
— subgen. Cinygmula: Kluge 1983b:16 (Abstract); 1988:305
— gen. Cinygmula McDunnough 1933: 75
— Cinygmula/g1: Kluge 2004: 192
Nominal taxon included:
Ororotsia/g
[g:1939]
References.
McDunnough 1933: 
*; –
Needham & Traver & Hsu 1935: *
*; – Tshernova 1974: *;
– 1976: *; – Edmunds
& Jensen & Berner 1976: *
*; – Kluge 1988: *;
– 1993: *; – 2004: *
* *.
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Autapomorphy of Cinygmula. (1) Larval caudalii have no swimming setae [see Pentamerotarsata (6)] (unlike Rhithrogena/fg3, where primary swimming setae are retained). When swimming, larvae of Cinygmula make the same dosoventral movements by abdomen, as larvae with developed swimming setae [see Rhithrogena/fg2 (1)]; thus, loss of primary swimming setae is not connected with adaptation. Non-unique apomorphy (see Index of characters [1.3.66]). Characters of Cinygmula of unclear phylogenetic status. (2) Fibrillose portions of tergalii [see Branchitergaliae (3)] are diminished or lost (unlike Rhithrogena/fg3, where they are always large). Non-unique apomorphy (see Index of characters [1.3.25]). Plesiomorphies of Cinygmula (unlike Rhithrogena/fg3). On 2nd+3rd segment of maxillary palp scraping setae have no denticles, occupy about 1/2 palp length only (Kluge 2004: Fig.62:A). Larval and adult patella-tibial suture is developed on middle and hind legs only (unlike larval Rhithrogena/fg3). Tergalii do not form sucking disc (while in some species tergalius I is markedly widened anteriorly). Costal and anal ribs of tergalii are situated on costal and anal margin correspondingly [see Branchitergaliae (3)] (only on tergalius I costal rib can be reduced). Subimago without non-concave pigmented area behind apex of lateroparapsidal suture (Kluge 2004: Fig.63:A). Penis has median titillators [see Rhithrogena/fg2 (4) and Pentamerotarsata (5)]. Variable characters of Cinygmula. Imaginal and subimaginal claws are usually ephemeropteroid, but in hutchinsoni [O.] – pointed (see Variable characters of Heptagennota and Index of characters [2.2.85]). Gonostyli usually have 2 apical segments, but in tetramera [C.] they have 3 apical segments. |
Size. Fore wing length 7–14 mm.
Distribution. Amphipacific Sector of Holarctic: Eastern Palaearctic and Nearctic; dominate in Eastern Palaearctic and Western Nearctic.
Nominal species in Cinygmula/g1:
adusta
Imanishi 1935 [Cinygma]
altaica Tshernova 1949 [Cinygmula]
— syn.subj.
cavum
[Cinygma]
andrianovae Tshernova & Belov 1982 [Cinygmula]
— syn.subj. malaisei [Cinygma]
atlantica McDunnough 1924 [Cinygma]
— syn.subj. subaequalis [Heptagenia]
autumnalis Tiunova
& Gorovaya 2012 [Cinygmula]
brunnea Tiunova 1990 [Cinygmula]
,,
cavum
Ulmer 1927 [Cinygma]
--/
confusa McDunnough
1924 [Cinygma]
— syn.subj. par [Cinygma]
dorsalis Imanishi 1935 [Cinygma]
gartrelli
McDunnough 1934 [Cinygmula]
grandifolia
Tshernova 1952 [Cinygmula]
— syn.subj. sapporensis [Siphlurus]
guentheri Braasch 1979 [Cinygmula]
— syn.subj. cavum
[Cinygma]
hirasana Imanishi 1935 [Cinygma]
--/
hutchinsoni Traver
1939 [Ororotsia]
— typus nominis Ororotsia
--/
hyalina McDunnough 1924 [Cinygma]
— syn.subj. par [Cinygma]
inermis Braasch
1983 [Cinygmula]
irina Tshernova & Belov 1982 [Cinygmula]
joosti Braasch 1977 [Cinygmula]
--/
kaszabi Landa & Soldán 1983 [Cinygmula]
— syn.subj. kurenzovi
[Cinygma]
kootenai McDunnough
1943 [Cinygmula]
kurenzovi
Bajkova 1965 [Cinygma]
--/
latifrons Tshernova & Belov 1982 [Cinygmula]
— syn.subj. hirasana [Cinygmula]
levanidovi Tshernova & Belov 1982 [Cinygmula]
--/
longissima
Ding &
Gong & Zhou 2024 [Cinygmula]
malaisei Ulmer 1927 [Cinygma]
mimus Eaton 1885 [Cinygma]
minuta
Braasch 1980 [Cinygmula]
oreophila Kustareva 1978 [Cinygmula]
— syn.subj. joosti [Cinygmula]
par Eaton
1885 [Cinygma]
,
-
picta Braasch & Soldán
1979 [Cinygmula]
— ?syn.subj. joosti [Cinygmula]
putoranica Kluge 1980 [Cinygmula]
--/
quadripunctata Braasch & Soldán 1980 [Cinygmula]
— ?syn.subj. joosti [Cinygmula]
ramaleyi Dodds 1923 [Ecdyonurus]
— typus nominis Cinygmula
reticulata
McDunnough 1934 [Cinygmula]
rougemonti Braasch & Soldán 1987 [Cinygmula]
— ?syn.subj. joosti [Cinygmula]
sapporensis Matsumura 1904 [Siphlurus]
--/
spinosa
Tshernova & Belov 1982 [Cinygmula]
— syn.subj. levanidovi [Cinygmula]
subaequalis Banks 1914 [Heptagenia]
tarda McDunnough 1929
[Cinygma]
tetramera
Tiunova & Gorovaya 2013 [Cinygmula]
tioga Mayo 1952 [Cinygmula]
tollandi Dodds 1923 [Iron]
— syn.subj. ramaleyi [Ecdyonurus]
unicolorata
Tshernova 1979 [Cinygmula]
uniformis
McDunnough 1934 [Cinygmula]
uyka Gorovaya & Tiunova 2013 [Cinygmula]
vernalis Imanishi 1935 [Cinygma]
zimmermanni
Braasch 1977 [Cinygmula]
— ?syn.subj. joosti [Cinygmula]
See also:
Radulapalpata
INCERTAE SEDIS
