CLADOENDESIS OF EPHEMEROPTERA

ABC

z   

Ameletus/fg2

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta
Ameletus/fg1 - Ameletus/fg2)

Nomen hierarchicum: Ameletus/fg2 [f:1991; g:1885] (sine Metreletus

In circumscription fits:

gen. Ameletus Eaton 1885: 210

Ameletus/fg2: Kluge 2004: 82

Nominal taxa included: 

Chimura/g [g:1915] 

Paleoameletus/g [g:1940]


References. Eaton 1883-1885: * *; Needham & Traver & Hsu 1935: * *; Edmunds & Jensen & Berner 1976: * *; Zloty 1996: * *; Kluge 2004: * * *; Kluge 2007: * * *.


Autapomorphies of Ameletus/fg2.

(1) Glossae [see Ameletus/fg1 (1)] are truncate, i.e. with widened straight apical margin; all apical setae form a regular apical-ventral row and are flat, widened toward apex and rounded apically (Kluge 2004: Fig.20:A). Unique apomorphy.

Characters of Ameletus/fg2 of unclear phylogenetic status.

(2) Larval protopenis has simple shape: each lobe is triangular or conic, with rounded apex and without incision between lateral lobe and ventral plate [which are present on imaginal penis see Ameletus/fg1 (6)]  (Kluge 2007: Fig.10, 21); apex of this larval penis lobe corresponds to the apex of imaginal lateral lobe. This shape of larval penis is retained not only in species whose imaginal lateral lobes are long, but also in oregonensis [Ameletus] whose imaginal lateral lobes are shorter than median plates and strongly divergent.

Plesiomorphies of Ameletus/fg2 (unlike Metreletus). Cubital field of fore wing without intercalaries, several (48) veins go from CuA to basitornal margin [see Anteritorna (1)]. Larval claws have no denticles.

Size. Fore wing length 717 mm.

Distribution. Holarctic and Oriental Region, dominate in Nearctic and Eastern Palaearctic; in Western Palaearctic occurs a single Holarctic species inopinatus [Ameletus].


Nominal species in Ameletus/fg2:

aequivocus McDunnough 1934 [Ameletus] syn.subj. sparsatus [Ameletus]

aetherea Navás 1915 [Chimura] typus nominis Chimura

alexandrae Brodsky 1930 [Ameletus] --/

alpinus Bengtsson 1913 [Ameletus] syn.subj. inopinatus [Ameletus]

altaicus Kluge 2007 [Ameletus] --/

alticolus McDunnough 1934 [Ameletus] syn.subj. celer [Ameletus]

amador Mayo 1939 [Ameletus]

andersoni Zloty 1996 [Ameletus]

 arlecchino Kluge 2007 [Ameletus montanus] --/

asiacentralis Soldán 1978 [Ameletus] syn.subj. alexandrae [Ameletus]

atratus Kang & Yang 1994 [Ameletus]

bellulus Zloty 1996 [Ameletus] -

browni McDunnough 1933 [Ameletus]

camtschaticus Ulmer 1927 [Ameletus] --/

cedrensis Sinitshenkova 1977 [Ameletus] --/

celer McDunnough 1934 [Ameletus] -

celeroides McDunnough 1934 [Ameletus] syn.subj. celer [Ameletus]

connectina McDunnough 1939 [Ameletus] syn.subj. velox [Ameletus]

connectus McDunnough 1936 [Ameletus] syn.subj. velox [Ameletus]

cooki McDunnough 1929 [Ameletus]

costalis Matsumura 1931 [Chimura] ,-,/

cristatus Bajkova 1976 [Ameletus]

crocerus Imanishi 1931 [Ameletus]

cryptostimulus Carle 1978 [Ameletus]

dissitus Eaton 1885 [Ameletus]

doddsianus Zloty 1996 [Ameletus]

dodecus Dubey 1971 [Ichthybotus]

edmundsi Zloty 1996 [Ameletus]

eugenii Sinitshenkova & Varykhanova 1989 [Ameletus] syn.subj. inopinatus [Ameletus]

exquisitus Eaton 1885 [Ameletus]

facilis Day 1952 [Ameletus] syn.subj. vancouvernensis [Ameletus]

falsus McDunnough 1938 [Ameletus]

formosus Kang & Yang 1994 [Ameletus]

imbellis Day 1952 [Ameletus]

inopinatus Eaton 1887 [Ameletus --/

janetae Kondratieff & Meyer 2010 [Ameletus

khasanensis Tiunova 2013 [Ameletus]

kyotensis Imanishi 1932 [Ameletus]

labiatus Sinitshenkova 1981 [Ameletus] --

lineatus Traver 1932 [Ameletus]

longulus Sinitshenkova 1981 [Ameletus] ,-,/

ludens Needham 1905 [Ameletus]

majusculus Zloty 1996 [Ameletus]

minimus Zloty & Harper 1999 [Ameletus]

monta Mayo 1952 [Ameletus] syn.subj. similor [Ameletus]

montanus Imanishi 1930 [Ameletus] --/

montivagus Kang & Yang 1994 [Ameletus]

oregonensis McDunnough 1933 [Ameletus] -

parvus Kluge 1979 [Ameletus] --/

pilatus Sinitshenkova 1981 [Ameletus] syn.subj. camtschaticus [Ameletus]

primitivus Traver 1939 [Ameletus] typus nominis Paleoameletus

pritchardi Zloty 1996 [Ameletus]

procerus Bajkova 1976 [Ameletus]

quadratus Zloty & Harper 1999 [Ameletus]

querulus McDunnough 1938 [Ameletus] syn.subj. shepherdi [Ameletus]

 rossicus Kluge 2007 [Ameletus montanus] --/

shepherdi Traver 1934 [Ameletus]  

similor McDunnough 1928 [Ameletus] /,

sparsatus McDunnough 1931 [Ameletus]

subalpinus Imanishi 1931 [Ameletus]

subnotatus Eaton 1885 [Ameletus] typus nominis Ameletus

suffusus McDunnough 1936 [Ameletus

tarteri Burrows 1987 [Ameletus]

tertius McDunnough 1938 [Ameletus]

tolae Zloty 1996 [Ameletus]

tuberculatus McDunnough 1939 [Ameletus] syn.subj. celer [Ameletus]

validus McDunnough 1923 [Ameletus]

vancouvernensis McDunnough 1933 [Ameletus]

velox Dodds 1923 [Ameletus] -

vernalis McDunnough 1924 [Ameletus]

walleyi Harper 1970 [Ameletus]


Examined also:

sp. (China) /