CLADOENDESIS OF EPHEMEROPTERA
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Leptophlebia/fg1 Atalophleboadentata Atalophlebopectinata Atalophleboculata Atalophlebomaxillata Atalophlebolinguata
... Choroterpes/fg1 - Thraulus/fg1)
Nomen hierarchicum: Thraulus/fg1 [g:1881]
In circumscription fits:
gen. Thraulus: ??
Nominal taxon included: Masharikella/g [g:1964]
References. Peters & Edmunds 1970: * *; Kang & Yang 1994: '.
Autapomorphies of Thraulus/fg1.
() Egg surface with unique relief in a form of net made by convex ridges: several (10?) sinuous ridges go from one pole to another, and neighbouring ridges are connected by cross-ridges in such a manner that cross-ridges located by two sides of each longitudinal ridge are regularly alternating (Kang & Yang 1994: Fig.1316). ??in Thraulus femoratus ??
() Each of two lamellae of tergalius [see Leptophlebia/fg1 (10)] is wide, oval, with numerous long slender marginal processes. Tergalii IIVII always have such structure; tergalius I either has the same structure (in hsui [Thraulus], mariae [Thraulus]), or its dorsal lamella is slender without processes (in torrentis [Masharikella]), or its both lamellae are slender without processes (in bellus [Thraulus], fatuus [Thraulus], macilentus [Thraulus], umbrosus [Thraulus] ...?), or tergalius consists of a single slender lamella without processes (in femoratus [Thraulus]). Tergalii with similar marginal processes are found in some other taxa of Leptophlebia/fg1 (see Index of characters [1.3.23]).
Characters of Thraulus/fg1 of unclear phylogenetic status.
() Larval patella-tibial suture is lost on all legs, while in imago and subimago it is retained on middle and hind legs, being absent on fore legs only (the same in Choroterpes/g1 - see Index of characters [1.2.18], ibid. [2.2.82]).
Plesiomorphies of Thraulus/fg1.
Imaginal and subimaginal claws of all legs are ephemeropteroid (see Index of characters [2.2.85]).
Size. Fore wing length 510 mm.
Distribution. Old World (Afrotropical, Oriental and Palaearctic Regions).
Nominal species in Thraulus/fg1:
bellus Eaton 1881 [Thraulus] typus nominis Thraulus ,-,-,-
bishopi Peters & Tsui 1972 [Thraulus]
demoulini Peters & Tsui 1973 [Thraulus]
fasciatus Kimmins 1956 [Hagenulus] typus nominis Masharikella
fatuus Kang & Yang 1994 [Thraulus]
femoratus Li & Liu & Zhou 2006 [Thraulus]
gopalani Grant & Sivaramakrishnan 1985 [Thraulus]
grandis Gose 1980 [Thraulus]
macilentus Kang & Yang 1994 [Thraulus]
maculatus Peters & Tsui 1972 [Thraulus] nom.praeocc. syn.obj. demoulini [Thraulus]
mudumalaiensis Soman 1991 [Thraulus]
semicastanea Gillies 1951 [Habrophlebiodes] ?
thraker Jacob 1988 [Thraulus]
torrentis Gillies (in Peters & Gillies & Edmunds) 1964 [Masharikella]
turbinatus Ulmer 1909 [Hagenulus]
umbrosus Kang & Yang 1994 [Thraulus]
sp. (Sulawesi, Mamasa) --
sp. (Sulawesi, Mamasa)
sp. (Papua, Baliem) /
sp. (Thailand) --/
Atalophlebolinguata INCERTAE SEDIS
Leptophlebia/fg1 INCERTAE SEDIS