CLADOENDESIS OF EPHEMEROPTERA

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Thraulodes

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Leptophlebia/fg1 Atalophleboadentata Atalophlebopectinata Atalophleboculata Atalophlebomaxillata Atalophlebolinguata 
... Hagenulus/fg1 Hermanellandria - Thraulodes)

Nomen hierarchicum: Thraulodes/g2 [g:1920] (sine Hermanella)

In circumscription fits:

 gen. Thraulodes Ulmer 1920: 33


References. Traver & Edmunds 1967; Kluge 2020: * * *


Autapomorphies of Thraulodes.

(1) Bases of gonostyli are brought together in a common unpaired gonostylar cavity of styliger; dorsal margin of styliger is projected caudally, and the gonostylar cavity is exposed ventrally-caudally (Kluge 2020: Figs 102, 110, 112).

(2) Telopenis [see Hermanellandria (1)] is tubular with a longitudinal cleft up to its base, i.e. represents a roll (in contrast to integral tube of Hermanellognatha and Simothraulopsis and non-tubular process of Farrodes) (Kluge 2020: Figs 38, 60, 103105).

(3) On hind tibia, stout setae of outer margin [see Leptophlebia/fg1 (8)] form two separated rows, outer-anterior and outer-posterior ones; long hairs (also initial for Leptophlebia/fg1) are situated between these two rows of stout setae and form one more row posteriad of them; row of recurved hairs is located on anterior side of tibia, more distally from inner margin than the inner-anterior row of stout setae, which is present on middle and hind legs (Kluge 2020: Figs 85, 89).

Characters of Thraulodes of unclear phylogenetic status. 

(4) Outer side of each mandible with distal setal bunch, but without middle setal bunch: long hairs forming row of field on outer margin of mandible, are denser and longer nearer the base of incisor (Kluge 2020: Figs 1718) (the same in some other taxa).

(5) Long setae of dorsal side of glossa form irregular row or stripe of a characteristic horseshoe shape (Kluge 2020: Fig. 20) (the same in Atopophlebia).

(6) Larva (whose body is flattened and widened, being adopted for inhabitancy under stones on current, as in many other Atalophlebolinguata) has unusual mode of swimming: it keeps legs laterally semi-bent in knees (as if sitting on substrate) and makes lateral swimming movements by abdomen (unlike most other Ephemeroptera, which make dorso-ventral swimming movements). The same in Kimminsula/g1 (see Index of characters [1b]).

Plesiomorphies of Thraulodes.

(7) Hind wing always retains furcations of RS and MP [furcation of MA is lost see Furcatergaliae (4)] (Kluge 2020: Fig. 41).

Size. Fore wing length 515 mm.

Distribution. Neotropical Region and south of Nearctic Region.


Nominal species in Thraulodes/g(1):

alapictus Lima & Mariano & Pinheiro 2013 [Thraulodes]

alboniger Kluge 2020 [Thraulodes] --/

amanda Mariano & Froehlich (in Mariano & Flowers & Froehlich) 2011 [Thraulodes]

arizonicus McDunnough 1942 [Thraulodes]

basimaculatus Giordano & Dominguez 2005 [Thraulodes]

bolivianus Dominguez 1986 [Thraulodes]  

bomplandi Esben-Petersen 1912 [Thraulus]  

bonito Goncalves & Da-Silva & Nessimian 2013 [Thraulodes]

borum [Thraulodes] nomen nudum

brunneus Koss 1966 [Thraulodes]  

calori Campos & Mariano 2019 [Thraulodes]

catoles Campos & Mariano 2019 [Thraulodes]

centralis Traver 1946 [Thraulodes]  

cochunaensis Dominguez 1987 [Thraulodes]  

consortis Dominguez 1987 [Thraulodes] ,

cryptodrilus Neito & Dominguez 2001 [Thraulodes]

daidaleus Thew 1960 [Thraulodes]  

eduardorum Medina & Perez 2010 [Thraulodes]  

eccentricus Lugo-Ortiz & McCafferty 1996 [Thraulodes] nomen dubium

ephippiatus Traver & Edmunds 1967 [Thraulodes]  

fascipennis Kluge 2020 [Thraulodes] --/

flavus Kluge 2020 [Thraulodes] --; /

flinti Dominguez 1987 [Thraulodes]  

furficulus Traver 1946 [Thraulodes]  

gonzalesi Traver & Edmunds 1967 [Thraulodes]  

grandis Lugo-Ortiz & McCafferty 1996 [Thraulodes] nomen dubium

guanare Chacon & Segnini & Dominguez 1999 [Thraulodes]  

hilaris Eaton 1892 [Thraulus]  

hilaroides Traver 1946 [Thraulodes]  

humeralis Navas 1935 [Thraulodes]  

insular Domínguez & Molineri & Zúñiga (in Zúñiga & Molineri & Domínguez & Cardona) 2015 [Thraulodes]

irretitus Navas 1924 [Thraulodes]  

itatiajanus Traver & Edmunds 1967 [Thraulodes]  

jones Gonçalves & Da-Silva & Nessimian 2010 [Thraulodes]  

latinus Cortes & Dias & Araújo 2021 [Thraulodes]

laetus Eaton 1884 [Thraulus] typus nominis Thraulodes  

lepidus Eaton 1884 [Thraulus]  /

limbatus Navas 1936 [Thraulodes]  

liminaris Dominguez 1987 [Thraulodes]  

ludmilae Kluge 2020 [Thraulodes] --

luisae Souto & Da-Silva & Nessimian 2014 [Thraulodes]

luizgonzagai Lima & Mariano & Pinheiro 2013 [Thraulodes]

lunatus Traver & Edmunds 1967 [Thraulodes]  

marianoi Silva & Salles & Pinto 2020 [Thraulodes]  syn.subj. basimaculatus [Thraulodes]

marreroi Chacon & Segnini & Dominguez 1999 [Thraulodes] --/

mexicanus Eaton 1884 [Thaulus]  

mucuy Chacon & Segnini & Dominguez 1999 [Thraulodes]  

nervosa Eaton 1892 [Choroterpes] ?

niger Kluge 2020 [Thraulodes] --/

nigrabdominalis Kluge 2020 [Thraulodes] -,

nigripes Kluge 2020 [Thraulodes]   /, --/

nigrotibialis Kluge 2020 [Thraulodes] --/

osiris Traver & Edmunds 1967 [Thraulodes]  

pacaya McCafferty & Baumbgardner & Guenther 2004 [Thraulodes] nomen dubium

packeri Traver & Edmunds 1967 [Thraulodes]  

panamensis Kluge 2020 [Thraulodes] --/

papilionis Traver & Edmunds 1967 [Thraulodes]  

paysandensis Traver 1964 [Thraulodes]

pedregoso Traver 1946 [Thraulodes]

pelicanus Mariano & Froehlich (in Mariano & Flowers & Froehlich) 2011 [Thraulodes]

pinga Souto & Da-Silva & Nessimian 2014 [Thraulodes]

pinhoi Lima & Mariano & Pinheiro 2013 [Thraulodes]

plicatus Needham & Murphy 1924 [Thraulodes]  syn.subj. telegraphicus [Thraulodes]

prolongatus Traver 1946 [Thraulodes]  

quevedoensis Flowers 2009 [Thraulodes] --/

regulus Traver & Edmunds 1967 [Thraulodes]  

rodrigoi Boldrini & Dantas & Lima 2018 [Thraulodes]

salinus Kilgore & Allen 1973 [Thraulodes] nomen dubium

schlingeri Traver & Edmunds 1967 [Thraulodes]   --/

sinuosus Mariano & Flowers (in Mariano & Flowers & Froehlich) 2011 [Thraulodes] --/

spangleri Traver & Edmunds 1967 [Thraulodes]  /, -

speciosus Traver 1934 [Thraulodes]  

sternimaculatus Lima & Mariano & Pinheiro 2013 [Thraulodes]

telegraphicus Needham & Murphy 1924 [Thraulodes]  --/

tenulineus Lugo-Ortiz & McCafferty 1996 [Thraulodes]  nomen dubium

traverae Thew 1960 [Thraulodes]  

trijuncta Banks 1918 [Thraulus]  

ulmeri Edmunds 1950 [Thraulodes]  

valens Eaton 1892 [Thraulus]  

venezuelana Ulmer 1943 [Thraulodes]  

viviparus Kluge 2020 [Thraulodes] -/,

xavantinensis Mariano & Froehlich (in Mariano & Flowers & Froehlich) 2011 [Thraulodes]

zonalis Traver & Edmunds 1967 [Thraulodes]  --/, /


Examined:

spp. (Panama) --/

spp. (Peru) --/


   

KEY TO IMAGINES

1(8) All tibiae either entirely dark (Kluge 2020: Figs 467469, 491492, 526527), or dark in proximal half (Kluge 2020: Fig. 437).

2(7) All femora uniformly dark brown (Kluge 2020: Figs 436437, 467469, 491492).

3(4) Fore wing with costal and subcostal fields contrastingly yellowish (Kluge 2020: Figs 427, 435). . . . niger Kluge 2020 [Thraulodes]

4(3) Fore wing with costal and subcostal fields not colored (Figs 465, 493).

5(6) Fore wing with dark macula on costal brace (Kluge 2020: Fig. 493). ♂: Abdominal terga IIVI light ocher, terga VIIX reddish (Kluge 2020: Fig. 495). . . . nigripes Kluge 2020 [Thraulodes]

6(5) Fore wing without dark macula on costal brace (Kluge 2020: Fig. 465). ♂: Abdominal terga unicolor (Kluge 2020: Fig. 466). . . . nigrabdominalis

7(2) All femora twice banded, i.e. with small light area at middle of femur between bands (Kluge 2020: Figs 526527). . . . nigrotibialis Kluge 2020 [Thraulodes]

8(1) At least middle and hind tibiae either entirely light (Kluge 2020: Fig. 2930), or light with dark apex (Kluge 2020: Figs 306307, 659660).

9(14) Fore tibia entirely dark brown (Fig. 597).

10(11) At least in male, proximal half of fore wing brown, distal half colorless (female unknown). . . . basimaculatus Giordano & Dominguez 2005 [Thraulodes] (= marianoi [Thraulodes])

11(10) No such difference between proximal and distal halves of fore wing.

12(13) All femora uniformly dark brown (Kluge 2020: Figs 597598). . . . lepidus Eaton 1884 [Thraulus]

13(12) Middle and hind femora yellowish with black bands in distal half. . . . venezuelana Ulmer 1943 [Thraulus]

14(9) Fore tibia at least partly light (Kluge 2020: Figs 28, 305, 339, 365, 574) (as well as middle and hind ones).

15(16) Fore tibia with brown band at middle, apex whitish; hind femur whitish with two narrow black bands close to apex (Kluge 2020: Figs 574575). . . . alboniger Kluge 2020 [Thraulodes]

16(15) Fore tibia with dark band adjacent to apex (Kluge 2020: Figs 28, 305, 339, 365). Hind femur not as above.

17(18) Femora of middle and hind legs yellowish, without dark maculae; only femur of fore leg with dark maculae apically. . . . papilionis Traver & Edmunds 1967 [Thraulodes]

18(17) Femora of middle and hind legs with dark maculae at least in distal half (Kluge 2020: Figs 2930, 97, 306307, 367).

19(22) Fore femur evenly blackish brown nearly all over its length (paler at extreme base and/or apex).

20(21) ♂: Penis without ears, medio-ventral ridge forms lapel projected laterad of lateral margin (Traver & Edmunds 1967: Fig. 67). . . . hilaris Eaton 1892 [Thraulus]

21(20) ♂: Penis with ears, no lapel projected laterad of lateral margin (Traver & Edmunds 1967: Fig. 51). . . . arizonicus McDunnough 1942 [Thraulodes]

22(19) Fore femur with darker maculae at least in distal half (Kluge 2020: Figs 28, 305, 339, 365, 398).

23(66) Middle and/or hind femora, besides dark band and/or maculae in distal half, with dark band or spot in proximal half (Kluge 2020: Figs 307, 341, 365367, 398401, 660)

24(57) Proximal half of middle and hind femora with band or wide spot (Kluge 2020: Figs 307, 341, 365367, 398401, 660).

25(26) Subcostal field of fore wing yellow, apically brown (Kluge 2020: Figs 365367). . . . flavus Kluge 2020 [Thraulodes]

26(25) Subcostal field of fore wing, if colored, with apex not darker than remainder part.

27(28) Fore wing with dark macula on furcation of MA larger than other maculae. . . . alapictus Lima, Mariano & Pinheiro 2013 [Thraulodes]

28(27) If dark maculae on fore wing present, macula on furcation of MA not larger than others (Kluge 2020: Fig. 398401).

29(30) Contrasting dark brown spot on whitish background on median furcasternal concavity of mesothorax (Kluge 2020: Fig. 400). . . . spangleri Traver & Edmunds 1967 [Thraulodes]

30(29) No such unpaired spot on median furcasternal concavity.

31(32, 33, 34) ♂: Penis with pair of apico-lateral projections directed proximally (Kimmins 1934: Fig. 8). . . . valens Eaton 1892 [Thraulus]

32(31, 33, 34) ♂: Penis with pair of lateral projections directed proximally (Campos & Mariano 2019: Figs 1012). . . . catoles Campos & Mariano 2019 [Thraulodes]

33(31, 32, 34) ♂: Penis with pair of lateral projections directed distally (Traver 1946: Fig. 30; Traver & Edmunds 1967: Fig. 65). . . . furficulus Traver 1946 [Thraulodes

34(31, 32, 33) ♂: Penis without additional lateral projections.

35(40) ♂: Penis in distal half parallel-sided, not widened apically (Kluge 2020: Fig. 310 or similar to Fig. 440).

36(37) ♂: Penis without ears (Traver & Edmunds 1967: Fig. 37). . . . ephippiatus Traver & Edmunds 1967 [Thraulodes]

37(36) ♂: Penis with ears directed caudally (Kluge 2020: Fig. 310; Traver & Edmunds 1967: Fig. 41).

38(39) ♂: Abdominal terga IIIX equally dark (Kluge 2020: Fig. 343). . . . zonalis Traver & Edmunds 1967 [Thraulodes]

39(38) ♂: Abdominal terga IIV contrastingly lighter than terga VIIX (Kluge 2020: Figs 298301). . . . fascipennis Kluge 2020 [Thraulodes]

40(35) ♂: Penis widened apically.

41(42) ♂: Penis hidden under very wide dorsal projection of styliger (Neito & Dominguez 2001: Figs 68). . . . cryptodrilus Neito & Dominguez 2001 [Thraulodes]

42(41) ♂: Penis not hidden, dorsal projection of styliger smaller.

43(50) ♂: Penis without lateral pouches.

44(45) ♂: Dorsal margin of styliger rounded, without median projection (Traver & Edmunds 1967: Fig. 63). . . . laetus Eaton 1884 [Thraulus]

45(44) ♂: Dorsal margin of styliger either pointed, or with median projection (Traver & Edmunds 1967: Fig. 44; Dominguez 1987: Figs 6, 8).

46(47) ♂: Abdominal terga IVVI with composite dark markings (Dominguez 1987: Figs 1920). . . . liminaris Dominguez 1987 [Thraulodes]

47(46) ♂: Abdominal terga IVVI with sublateral dark spots (Dominguez 1987: Figs 1516).

48(49) Distributed in South America; genitalia as figured by Dominguez (1987: Fig. 6). . . . consortis Dominguez 1987

49(48) Distributed in North America; genitalia as figured by Traver & Edmunds (1967: Fig. 44). . . . speciosus Traver 1934 [Thraulodes]

50(43) ♂: Penis with lateral pouches.

51(52, 53, 54, 55, 56) ♂: Abdominal coloration and genitalia as described by Traver & Edmunds (1967: Figs 89, 31). . . . packeri Traver & Edmunds 1967 [Thraulodes]

52(51, 53, 54, 55, 56) ♂: Abdominal coloration and genitalia as described by Traver & Edmunds 1967: Fig. 26). . . . . daidaleus Thew 1960 [Thraulodes]

53(51, 52, 54, 55, 56) ♂: Abdominal coloration and genitalia as described by Traver & Edmunds (1967: Figs 45, 46). . . . paysandensis Traver 1964 [Thraulodes]

54(51, 52, 53, 55, 56) ♂: Abdominal coloration and genitalia as described by Traver & Edmunds (1967: Figs 23, 66). . . . . traverae Thew 1960 [Thraulodes]

55(51, 52, 53, 54, 56) ♂: Abdominal coloration and genitalia as described by Lima et al. (2013: Figs 1011, 1820). . . . luizgonzagai Lima, Mariano & Pinheiro 2013 [Thraulodes]

56(51, 52, 53, 54, 55) ♂: Abdominal coloration and genitalia as described by Lima et al. (2013: Figs 1617, 3132). . . . pinhoi Lima, Mariano & Pinheiro 2013 [Thraulodes]

57(24) Proximal half of middle and hind femora with small spot (besides band in distal half).

58(61) ♂: Abdominal terga IIIX equally dark.

59(60) ♂: Penis with lateral pouches wider than apex (Mariano et al. 2011: Figs 910). . . . amanda Mariano & Froehlich (in Mariano, Flowers & Froehlich) 2011 [Thraulodes]

60(59) ♂: Penis with lateral pouches narrower than apex (Mariano et al. 2011: Figs 43). . . . xavantinensis Mariano & Froehlich (in Mariano, Flowers & Froehlich) 2011 [Thraulodes]

61(58) ♂: Abdominal terga IIV contrastingly lighter than terga VIIX.

62(63) ♂: Dorsal margin of styliger rounded, without median projection (Lima et al. 2013: Fig. 12). . . . sternimaculatus Lima, Mariano & Pinheiro 2013 [Thraulodes]

63(62) ♂: Dorsal margin of styliger with median projection.

64(65) ♂: Abdominal terga IIVI with a pair of dark submedian spots (besides pair of midway spots) (Souto et al. 2014: Figs 14). . . . luisae Souto, Da-Silva & Nessimian 2014 [Thraulodes] and pinga Souto, Da-Silva & Nessimian 2014 [Thraulodes]

65(64) ♂: Abdominal terga without submedian spots. . . . insular Domínguez, Molineri & Zúñiga (in Zúñiga, Molineri, Domínguez & Cardona) 2015 [Thraulodes] and eduardorum Medina & Perez 2010 [Thraulodes]

66(23) Middle and hind femora light, with dark macula(e) only in distal part (Kluge 2020: Figs 30, 97).

67(68) Hind femur without distinct bands, darker toward apex. ♂: penis unusually sharply widened, with lateral denticles (Traver 1946: Fig. 27; Traver & Edmunds 1967: Fig. 49). . . . prolongatus Traver 1946 [Thraulodes]

68(67) Hind femur with brown or reddish apical band usually bordered by darker brown on proximal edge (Kluge 2020: Figs 30, 97). ♂: Penis not as above.

69(70) ♂: All abdominal terga IX dark brown. Dorsal margin of styliger rounded, without median projection; spear-like rolls very long and thick, directed caudally (Traver & Edmunds 1967: Fig. 42). . . . regulus Traver & Edmunds 1967 [Thraulodes]

70(69) ♂: At least abdominal terga IIV light.

71(72) ♂: Abdominal terga IIVI with grayish brown dome-shaped patches (Traver & Edmunds 1967: Figs 1011). . . . gonzalesi Traver & Edmunds 1967 [Thraulodes]

72(71) ♂: Abdominal coloration not as above.

73(80) ♂: Abdominal terga IIV mostly light, VIVII mostly colored (Kluge 2020: Figs 198, 232, 264).

74(75) ♂: Penis long and parallel-sided (Dominguez 1987: Fig. 10). . . . cochunaensis Dominguez 1987 [Thraulodes]

75(74) ♂: Penis widened apically (Kluge 2020: Figs 204, 242, 268269).

76(77) ♂: Spear-like rolls directed dorsally (Kluge 2020: Fig. 267). . . . quevedoensis Flowers 2009 [Thraulodes]

77(76) ♂: Spear-like rolls directed ventrally (Kluge 2020: Figs 203, 240).

78(79) ♂: Apices of penis lobes as wide as gonostylar cavity of styliger (Kluge 2020: Fig. 242). Distribution: from Panama to Venezuela. . . . marreroi Chacon & Segnini & Dominguez 1999 [Thraulodes]

79(78) ♂: Apices of penis lobes wider than gonostylar cavity of styliger (Kluge 2020: Fig. 204). Distribution: Peru and Brazil. . . . schlingeri Traver & Edmunds 1967 [Thraulodes]

80(73) ♂: Abdominal terga IIVI mostly light, VII mostly colored (Kluge 2020: Figs 25, 52, 99, 136, 166).

81(84) ♂: Spear-like rolls directed ventrally-proximally, with groove widely opened (Kluge 2020: Figs 38, 60).

82(83) ♂: Spear-like rolls S-shaped; dorsal extension of styliger wide and not separated from styliger (Kluge 2020: Fig. 60). . . . sinuosus Mariano & Flowers (in Mariano & Flowers & Froehlich 2011) [Thraulodes]

83(82) ♂: Spear-like rolls very short, lanceolate; dorsal extension of styliger finger-like and separated from styliger (Kluge 2020: Figs 38, 40). . . . ludmilae Kluge 2020 [Thraulodes]

84(81) ♂: Spear-like rolls directed medially (medially, medially-ventrally, or medially-dorsally) (Kluge 2020: Figs 112120, 145, 169).

85(86) ♂: Penis lobes narrowed apically (Gonçalves et al. 2010: Fig. 5). . . . jones Gonçalves & Da-Silva & Nessimian 2010 [Thraulodes]

86(85) ♂: Penis lobes not narrowed apically (Kluge 2020: Figs 112120, 145, 169).

87(90) ♂: Abdominal sterna IIVII with 2 or 3 pairs of dark dots.

88(89) ♂: Penis lobes long, equally narrow all over length (Traver & Edmunds 1967: Fig. 43). . . . itatiajanus Traver & Edmunds 1967 [Thraulodes]

89(88) ♂: Penis lobes short and greatly expanded laterally at apex (Dominguez 1987: Fig. 7). . . . flinti Dominguez 1987 [Thraulodes]

90(87) ♂: Abdominal sterna IIVII with no more than one pair of dots.

91(94) ♂: Penis without lateral pouches (Kluge 2020: Figs 112120).

92(93) ♂: Penis lobes short and sharply widened apically (Boldrini et al. 2018: Figs 67). Abdominal terga IIVI with reddish brown band adjacent to posterior margin (ibid: figs 23). . . . rodrigoi Boldrini & Dantas & Lima 2018 [Thraulodes]

93(92) ♂: Penis lobes longer and slightly widened apically (Kluge 2020: Figs 112120). Abdominal terga IIVI without contrasting dark band (Kluge 2020: Figs 97, 99). . . . telegraphicus Needham & Murphy 1924 [Thraulodes]

94(91) ♂: Penis with lateral pouches (Kluge 2020: Figs 144, 171).

95(101) ♂: Lateral pouches large, so that at midlength penis as wide or wider than at apex.

96(97) ♂: Dorsal margin of styliger rounded, without median projection (Chacon et al. 1999: Fig. 5). . . . mucuy Chacon & Segnini & Dominguez 1999 [Thraulodes]

97(96) ♂: Dorsal margin of styliger with median projection.

98(99, 100) ♂: Abdominal coloration and genitalia as described by Traver & Edmunds (1967: Fig. 33). . . . ulmeri Edmunds 1950 [Thraulodes]

99(98, 100) ♂: Abdominal coloration and genitalia as described by Mariano et al. (2011: Figs 1415). . . . pelicanus Mariano & Froehlich (in Mariano & Flowers & Froehlich) 2011 [Thraulodes]

100(98, 99) ♂:Abdominal coloration and genitalia as described by Gonçalves et al. (2013: Figs 46). . . . bonito Gonçalves & Da-Silva & Nessimian 2013 [Thraulodes]

101(95) ♂: Lateral pouches small, so that at midlength penis much narrower than at apex (Kluge 2020: Figs 144, 171).

102(103) ♂: Abdominal terga VIIIX dark reddish brown, tergum VII lighter. . . . centralis Traver 1946 [Thraulodes]

103(102) ♂: Abdominal tergum VII not lighter than tergum VIII (Kluge 2020: Figs 136, 166).

104(105, 106) Fore femur, besides dark band in distal half without dark marking in proximal half. . . . calori Campos & Mariano 2019 [Thraulodes]

105(104, 110) Fore femur, besides dark band in distal half, with dark band in proximal half. . . . trijuncta Banks 1918 [Thraulus] and bolivianus Dominguez 1986 [Thraulodes]

106(104, 115) Fore femur, besides dark band in distal half, with small dark spot in proximal half (Kluge 2020: Figs 134, 160).

107(108) ♂: Pronotum grayish-brown with white spots. . . . guanare Chacon, Segnini & Dominguez 1999 [Thraulodes]

108(107) ♂: Pronotum white (Kluge 2020: Figs 134, 136, 157).

109(110) Macula on costal brace not continuous posteriorly (Kluge 2020: Figs 134136). Scutellum white (Kluge 2020: Fig. 136). . . . panamensis Kluge 2020 [Thraulodes]

110(109) Macula on costal brace continuous posteriorly (Kluge 2020: Figs 157159). Scutellum brown (Kluge 2020: Fig. 159). . . . viviparus Kluge 2020 [Thraulodes]