CLADOENDESIS OF EPHEMEROPTERA

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Hagenulus/fg2

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna Bidentiseta Furcatergaliae  
Leptophlebia/fg1 Atalophleboadentata Atalophlebopectinata Atalophleboculata Atalophlebomaxillata Atalophlebolinguata 
... Hagenulus/fg1 - Hagenulus/fg2)

Nomen hierarchicum: Hagenulus/fg2 [f:1994; g:1882] (sine Hermanella, Thraulodes, Ulmeritus, Miroculis; incl. ) 

In circumscription fits:

 gen. Hagenulus: Kluge 1994b: 255

 Hagenulus/fg2: Kluge 2008: 387


References. Kluge 1994: .


Autapomorphies of Hagenulus/fg2.

() Apical-ventral row of pectinate setae [see Leptophlebia/fg1 (1)] is interrupted in middle forming two separate portions median portion (710 setae) and lateral portion (715 setae); these portions do not form common line: most median bristles of the lateral portion are smaller and arise more distally than most lateral bristles of the median portion (Figs#). In other Leptophlebia/fg1 this row is either integral and straight, or has curvation in middle (see Index of characters [1.1.31]).

() Penis with a pair of small pointed processes arising from ventral side near middle and usually directed medially; these processes are developed either both in subimago and imago, or in subimago only (in (in Hagenulus/fg3 morrisonae [H.] and Turquinophlebia/g1 grandis [H.]). In Careospina and Traverina these processes are completely absent, probably secondarily reduced.

Characters of Hagenulus/fg2 of unclear phylogenetic status. 

????() On fore wing fork of MP is symmetrical. Non-unique apomorphy (see Index of characters [2.2.46]).

() In cubital field of fore wing most anterior of two intercalaries [see Leptophlebia/fg1 (7)] is basally connected with CuP (see Index of characters [2.2.51]).

Plesiomorphies of Hagenulus/fg2.

...

Size. Fore wing length ?? mm.

Age and distribution. Cenozoic recent; Neotropical Region, dominate in Caribean area.


The taxon Hagenulus/fg2 is divided into:

1. Hagenulus/fg3

2. Borinquena

3. Turquinophlebia

4. Poecilophlebia

5. Careospina

6. Traverina

7. Hagenulopsis

8. Neohagenulus

???Dominican amber