CLADOENDESIS OF EPHEMEROPTERA
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Rhithrocloeon/fg4

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata 
pm.Protopatellata Rhithrocloeon/fg1 Rhithrocloeon/fg2 Rhithrocloeon/fg3 - Rhithrocloeon/fg4

Nomen hierarchicum: Rhithrocloeon/fg4 [g:1985] (sine Kivuiops)

In circumscription fits:

— gen. Rhithrocloeon: Lugo-Ortiz & McCafferty 1996

— Rhithrocloeon/fg3: Kluge 2012

— Rhithrocloeon/fg4: Kluge 2015

References. Gillies 1988: *; – Kluge 2012:1-13 (as Rhithrocloeon/fg3): *

Autapomorphy of Rhithrocloeon/fg4.

(1) Prostheca of right mandible is slender, seta-like, without denticles (Gillies 1988: Figs 34, 36) (unlike prostheca of left mandible, which is wide and dentate, as in all Rhithrocloeon/fg1). This differs both from Kivuiops (whose right prostheca is widened) and from Bugilliesia (whose right prostheca is plesiomorphically moderately narrow and dentate (Gillies 1990: Figs 71–73)) (see Index of characters [1.1.25]).

Character of Rhithrocloeon/fg4 of unclear phylogenetic status. 

(2) Labrum has uniform structure in both species: widened distally, with median incision small; in the specimen examined, dorsal surface with 1 pair of latero-distal setae and a pair of submedian setae (Kluge 2012: Fig.24).

Plesiomorphy of Rhithrocloeon/fg4.

(3) Unlike Kivuiops, maxilla [of «Cloeon-type» — see Rhithrocloeon/fg2 (3)] retains more than one seta in inner-dorsal row.

Size. Fore wing length 5–7 mm (see Tetramerotarsata).

Distribution. Afrotropical Region.

Nominal species in Rhithrocloeon/fg4:

indicator Gillies 1985 [Rhithrocloeon] -

permirum Kopelke 1980 [Cloeon] — typus nominis Rhithrocloeon --/

Examined also:

sp. (Tanzania): Gillies 1988:56; sp.1: Kluge 2012:1-13

sp.2 (Uganda): Kluge 2012:1-13

See also:

Turbanoculata INCERTAE SEDIS