CLADOENDESIS OF EPHEMEROPTERA
Tetramerotarsata Liberevenata Turbanoculata pm.Protopatellata
Rhithrocloeon/fg1 Rhithrocloeon/fg2 - Rhithrocloeon/fg3
Nomen hierarchicum: Rhithrocloeon/fg3 [g:1985] (sine Mutelocloeon, Bugilliesia; incl. Kivuiops)
In circumscription fits:
gen. Rhithrocloeon Gillies 1985
Rhithrocloeon/fg2: Kluge 2012
Rhithrocloeon/fg3: Kluge 2015
References. Gillies 1985: ; 1988: ; Kluge 2012:1-13 (as Rhithrocloeon/fg2): * *; Kluge 2015
Autapomorphies of Rhithrocloeon/fg3.
(1) Apical setae of paraglossa are shifted to ventral side, so that apical margin is free from setae (Kluge 2012: Fig.10) (unique apomorphy); these setae form 3 regular rows. Shape of labium is characteristic and identical in all species: paraglossa is widened apically; labial palp has 2nd segment bluntly projected medially (unlike sharply projected in Bugilliesia).
(2) Larval claw [slender see Rhithrocloeon/fg2 (4)] has only one row of denticles (Kluge 2012: Fig.7) (instead of initial two rows). The same in some other taxa of Protopatellata and in Anteropatellata-Baetungulata (see Index of characters [1.2.21]).
Characters of Rhithrocloeon/fg3 of unclear phylogenetic status.
(3) Hind wings absent; larva has no vestiges of hind protoptera (non-unique apomorphy see Index of characters [2.2.59]). Among Rhithrocloeon/fg1, the same in Adnoptilum; Mutelocloeon has vestigial hind wings; possibly, synapomorphy.
(4) Each tergalius IVII is bordered by integral marginal rib, whose whole distal part (including apex of tergalius) bears a regular row of small oblique seta-bearing denticles. The same in many other taxa, but not in Bugilliesia, whose tergalii have apex soft and non-denticulate. In Rhithrocloeon/fg2 shape of tergalii individually varies from oval to lanceolate, symmetric or slightly asymmetric (Gillies 1988: Figs 4042).
Size. Fore wing length 37 mm (see Tetramerotarsata).
Distribution. Afrotropical Region.
|The taxon Rhithrocloeon/fg3 is divided into:|
Turbanoculata INCERTAE SEDIS