CLADOENDESIS OF EPHEMEROPTERA
Tetramerotarsata Liberevenata Turbanoculata - pm.Protopatellata)
Nomen hierarchicum: Afroptilum/f1=Centroptiloides/g1 [f:1997; g:1918]
Nomen circumscribens: Protopatellata Kluge & Novikova 2011
In circumscription fits:
— subfam. Afroptilinae Kluge 1997: 533
— plesiomorphon Afroptilum/f1=Centroptiloides/g1: Kluge 2004: 105
— plesiomorphon Protopatellata Kluge & Novikova 2011: 5
References. Kluge 1997c: * * (plesiomorphies are wrongly called "Apomorphies"); – Kluge 2004: *; – Kluge & Novikova 2011: * * *; – Kluge 2015
Diagnostic plesiomorphiy of Protopatellata. In both sexes, larval, subimaginal and imaginal patella-tibial suture is present on middle and hind legs only. In some representatives the patella-tibial suture is lost on all legs, but is never present on fore legs (Kluge 2012: Fig.5-6) (unlike most larval and female Anteropatellata – see Index of characters [1.2.18] and Index of characters [2.2.]).
Other plesiomorphies of Protopatellata.
Setae on inner margin of mandibles [see Turbanoculata (6)] are usually present.
In subimago of both sexes at least most distal tarsomere of each leg is covered with pointed microlepides. The same in some Anteropatellata (see Index of characters [2.2.75], [220.127.116.11], [2.2.83]).
Subimaginal gonostyli buds under cuticle of mature larva are curved by the "Afroptilum-Cloeon-type" [see Turbanoculata (13)]: 2nd (the longest) segment is directed leterally. 3rd (distalmost) segment can be also directed laterally ("Afroptilum-type") (in sudafricanum [Centroptilum]), or is directed caudally ("Cloeon-type") (see Index of characters [2.3.9]); in Indocloeon folded gonostyli buds look unusually due to modification of its gonostyli [see below, Indocloeon (1)]. Exceptioon is made only by Rhithrocloeon/fg1, whose gonostyli and their buds have peculiar structure [see below, Rhithrocloeon/fg1 (1.2)].
On fore wing in each space only one marginal intercalary can be present [see Liberevenata (2)].
Size. Fore wing length 4–12 mm (see Tetramerotarsata).
Distribution. Mainly Afrotropical Region; also Oriental and Holarctic Regions.
|The plesiomorphon Protopatellata (or Afroptilum/f1=Centroptiloides/g1) is divided into:|
.... Indocloeon/g1 (incl. Hindocloeon)
.... Crassabwa/g1 (incl. Susua)
Combines characters of Protopatellata and Anteropatellata-Cloeon/fg1: Anafroptilum
Revision of these taxa is in progress