CLADOENDESIS OF EPHEMEROPTERA

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Nigrobaetis/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata
 Anteropatellata Baetovectata Baetungulata - Nigrobaetis/g1)

Nomen hierarchicum: Nigrobaetis/g1 [g:1987] (incl. Takobia)

In circumscription fits:

— subgen. Nigrobaetis: Novikova & Kluge 1987: 8;

— gen. Nigrobaetis: Kazlauskas 1972: 338 (nomen nudum).


References. Novikova & Kluge 1987: * *; – Kluge 1994: * *; – Kluge & Novikova 2014: * * *


Autapomorphies of Nigrobaetis/g1.

...

Characters of Nigrobaetis/g1 of unclear phylogenetic status. 

() Hind wing [see Turbanoculata (10)], if present, of the "Nigrobaetis-type": not narrowed, with non-curved pointed costal projection, with three longitudinal veins among which the second is bifurcate (i.e. MA arises from RA). The same in some other taxa (see Index of characters [2.2.60]). In some Oriental species hind wings are lost {?minutus Müller-Liebenau 1984 [Baetis]}.

() Larval antennae bases are turned together, frons between them forms a longitudinal keel (Kluge 1997: Tab.10:10; Lugo-Ortiz & de Moor 2000: Fig.1). Non-unique character, the same in some other taxa (Fallceon et al.), partly in Labiobaetis/f2=g1 and in Paracloeodes (see Index of characters [1.1.3]).

() Mandible ?? with setae between kinetodontium and mola.

() Larval abdominal terga with scales in operculate sockets (Kluge 1997: Tab.10:11) [see Turbanoculata (12)] (see Index of characters [1.3.4]).

Plesiomorphies of Nigrobaetis/g1.

In mature larva ready to moult to subimago, buds of subimaginal gonostyli are folded under larval cuticle by the "Nigrobaetis-type": that 2nd segment of gonostylus is directed posteriorly and curved by its convexity medially and by apex laterally (Kluge & Novikova 2014: Fig.76) [see Baetovectata (3)] (see Index of characters [2.3.9]). 

Larval paracercus is always well-developed.

Variable characters of Nigrobaetis/g1. In some species (digitatus [B.] and acinaciger [B.]) tergalii VII or VI–VII are crescent-shaped (Kluge 1997: Tab.10:8-9) (the same in Acerpenna); in other species all tergalii have usual oval shape.

Size. Fore wing length 3–8 mm (see Tetramerotarsata).

Distribution. Arctogea (Holarctic, Oriental and Afrotropical Regions).


The taxon Nigrobaetis/g1 is divided into:

1. plesiomorphon Nigrobaetis/g2

2. Takobia/g1