Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata 
 pm.Protopatellata Rhithrocloeon/fg1 Rhithrocloeon/fg2 Rhithrocloeon/fg3 - Kivuiops

Nomen hierarchicum: Kivuiops/g(1) [2007]

In circumscription fits:

— gen. Kivua Lugo-Ortiz & McCafferty 1996

— gen. Kivuiops Lugo-Ortiz & McCafferty 2007

— gen. Novokivua Özdikmen 2008

Objective synonyms of Kivuiops/g:

Kivua/g [g:1996] nom.praeocc.

Novokivua/g [2008] 

References. Kluge 2012:1-13: * *

Autapomorphies of Kivuiops.

(1) Prostheca of right mandible is widened, stout and dentate, similar to prostheca of left mandible (Kluge 2012: Fig.30). This differs both from Rhithrocloeon/fg3 (whose right prostheca is seta-like [comp.: Rhithrocloeon/fg4 (1)] and from Bugilliesia (whose right prostheca is plesiomorphically narrow and dentate (Gillies 1990: Figs 71–73)).

(2) On maxilla [of «Cloeon-type» – see Rhithrocloeon/fg2 (3)], inner-dorsal row of setae (which represents continuation of the row of dentisetae in proximal direction) consists of a single stout seta, which is especially long (much longer than dentisetae) and bent proximally (Kluge 2012: Fig.27); rarely one smaller seta locates proximad to it. Unlike Kivuips, in Rhithrocloeon/fg3 and Bugilliesia the inner-dorsal row retains several setae, which are not so long and not bent proximally.

(3) On labium [modified – see Rhithrocloeon/fg3 (1)] setae on inner side of glossae are short and stout (Kluge 2012: Fig.10) (unlike Rhithrocloeon/fg4, whose setae are longer and thinner – Kluge 2012: Fig.11).

(4) Larval paraproct has numerous small fine spines, directed perpendicular to its margin (Kluge 2012: Fig.36) (unlike Rhithrocloeon/fg4, whose paraproct has fewer oblique spines – Kluge 2012: Fig.37)

(5) On cuticle of subimaginal wing each microtrichion arises from center of a large brown round spot on colorless background (Kluge 2012: Fig.22). Examined species of Bugilliesia have no such spots. In Rhithrocloeon/fg4 permirum [Cloeon] this character ois absent (examined in 2016).

Characters of Kivuiops of unclear phylogenetic status. 

(6) All species have similar shape of gonostyli (Kluge 2012: Fig.12-18, 20, 31), different from gonostyli of each two species of Rhithrocloeon/fg3. Possibly, plesiomorphy.

Size. Fore wing length 3–6 mm (see Tetramerotarsata).

Distribution. Afrotropical Region.

Nominal species in Kivuiops/g(1):

elgonensis Kluge 2012 [Rhithrocloeon (Kivuiops)] --, //

elouardi Gillies 1989 [Rhithrocloeon]

insuetum Kopelke 1980 [Cloeon] — typus nominis Kivuiops --/

munyagae Kluge 2012 [Rhithrocloeon (Kivuiops)] -/, //

Examined also:


See also:


Turbanoculata INCERTAE SEDIS