CLADOENDESIS OF EPHEMEROPTERA

ABC

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Heterocloeon

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata
Liberevenata Turbanoculata Anteropatellata Baetovectata Baetungulata Baetofemorata - Heterocloeon)

Nomen hierarchicum: Heterocloeon/g1 [g:1925] (incl. Rheobaetis

In circumscription fits:

 gen. Heterocloeon McDunnough 1925: 175

 gen. Rheobaetis Müller-Liebenau 1974: 555

Nominal taxon included: 

Rheobaetis/g [g:1974]

Iswaeon/g [2005]

Jubilatum/g [2005]


References. Needham & Traver & Hsu 1935: * *; Müller-Liebenau 1974 (Rheobaetis): * *; Edmunds & Jensen & Berner 1976: * *; ...?


Autapomorphies of Heterocloeon.

(1) Claw with second row of denticles or at least a ridge on its place (McCafferty, Waltz, Webb & Jacobus 2005). The same secondary second row of denticles independently appeared in Andesiops/g1 (incl. Deceptiviosa); primary second row is present in Cloeon/fg1, plesiomorphon Afroptilum/f1=Centroptiloides/g2 and other taxa not belonging to Baetungulata.

() Labial palp has 2nd+3rd segments long and parallel-sided (muscle in 2nd segment is retained).

Characters of Heterocloeon of unclear phylogenetic status. 

() Imaginal and subimaginal anteronotal protuberance of mesonotum is sharply projected (Waltz 1994) (see Index of characters [2.2.5]). Possibly, synapomorphy with Acentrella/fg1 and/or Plauditus.

() Larva with gills in bases of fore legs (the same in some other Turbanoculata see Index of characters [1d]).

() On larval claw, denticles of inner-posterior row are distinctly developed [being much shorter than denticles of inner-anterior row see Baetungulata (1)] . ...?

Plesiomorphies of Heterocloeon.

Tibia without longitudinal row of setae (unlike Acentrella/fg1)

Size. Fore wing length 47 mm (see Tetramerotarsata).

Distribution. Nearctic.


Nominal species in Heterocloeon/g1:

anoka Daggy 1945 [Pseudocloeon]  typus nominis Iswaeon

berneri Müller-Liebenau 1974 [Rheobaetis]  

curiosum McDunnough 1923 [Centroptilum]  typus nominis Heterocloeon  

edmundsi Jensen 1969 [Pseudocloeon]  

frivolus McDunnough 1925 [Baetis]  

petersi Müller-Liebenau 1974 [Rheobaetis]  typus nominis Rheobaetis

traverae Müller-Liebenau 1974 [Rheobaetis] syn.subj. curiosum [Centroptilum]